# file: DRB4_nuc.txt
# date: 2025-04-09
# version: IPD-IMGT/HLA 3.60.0
# origin: https://github.com/ANHIG/IMGTHLA/blob/Latest/alignments/DRB4_nuc.txt
# repository: https://raw.githubusercontent.com/ANHIG/IMGTHLA/Latest/alignments/DRB4_nuc.txt
# author: IPD Team (ipdsubs@anthonynolan.org)
                   
 cDNA              1
 AA codon          -29
                   |
 DRB1*01:01:01:01  ATG GTG TGT CTG AAG CTC CCT GGA GGC TCC TGC ATG ACA GCG CTG ACA GTG ACA CTG ATG GTG CTG AGC TCC CCA 
 DRB4*01:01:01:01  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:01:03  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:01:04  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:01:05  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:01:06  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:02     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:03     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:04     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:05     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:06     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:07     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:08     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:09     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:10     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:01:11     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:12     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:13     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:01:14     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:02:01:01  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:02:01:02  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:01  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:02N --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:03  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:04  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:05  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:06  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:07  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:08  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:09  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:10  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:11  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:12  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:13N --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:14  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:15  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:16  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:17  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:18  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:19  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:20  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:21  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:22  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:23  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:24  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:25  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:26N --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:27  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:28N --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:29  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:01:30  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:01  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:02  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:03  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:04  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:05  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:06  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:07  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:08  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:09  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:02:10  --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:03     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:04     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:05     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:06     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:07     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:08     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:09     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:10     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:11     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:12     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:13     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:14     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:15     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:16     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:17     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:18     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:19     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:20     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:21     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:22     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:23     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:24     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:25     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:26     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:27     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:28     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:29     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:30     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:31     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:32     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:33     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:34     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:03:35     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:36     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:03:37     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:04        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:05        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:06        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:07:01     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:07:02     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:08        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:09        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:10        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:11        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:12        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:13        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:14N       --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:15        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:16N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:17        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:18        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:19        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:20        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:21        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:22        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:23        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:24        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:25        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:26        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:27        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:28        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:29        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:30        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:31        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:32        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:33        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:34        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:35        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:36        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:37        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:38N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:39        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:40        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:41        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:42        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:43        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:44        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:45        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:46        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:47        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:48        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:49        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:50        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:51        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:52        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:53        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:54:01N    *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:54:02N    *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:55        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:56N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:57N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:58        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:59        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:60        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:61:01N    *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:61:02N    *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:62        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:63        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:64        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:65N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:66        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:67        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:68        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:69        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:70        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:71N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:72:01     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:72:02     --- --- --- --- --- --- --- --- --- --- --T --- G-- --- --- --- --- --- T-- -C- --- --- --- --- --- 
 DRB4*01:73        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:74        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:75        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:76:01     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:76:02     *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:77        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:78        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:79        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:80N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:81        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:82        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:83        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:84N       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:85        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:86        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:87        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:88        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:89        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:90        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:91        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:92        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:93        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:94        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:95        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:96        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:97        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:98        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:99        *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:100       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:101       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:102       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:103       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:104       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:105       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:106       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:107       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:108N      *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:109       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:110       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:111       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:112       *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
 DRB4*01:113N      *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 
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 cDNA              76
 AA codon          -4
                   |
 DRB1*01:01:01:01  CTG GCT TTG GCT GGG GAC ACC CGA C|CA CGT TTC TTG TGG CAG CTT AAG TTT GAA TGT CAT TTC TTC AAT GGG ACG 
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 DRB4*01:03:28     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:03:29     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:03:30     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:03:31     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:26        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G -A- --- --- C-- --- --- --- 
 DRB4*01:27        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:28        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:41        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- -T- 
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 DRB4*01:45        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:49        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:52        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G G-- --- --- C-- --- --- --- 
 DRB4*01:53        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:54:01N    *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:54:02N    *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:55        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:56N       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:57N       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:58        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:59        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:60        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:61:01N    *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:61:02N    *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:62        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:63        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:64        *** *** *** *** *** *** *** *** *|-- -A- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:65N       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:66        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- A-G --- --- --- C-- --- --- --- 
 DRB4*01:67        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:68        *** *** *** *** *** *** *** *** *|-- -T- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:69        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- -C- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:70        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:71N       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:72:01     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:72:02     --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:73        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:74        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:75        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:76:01     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:76:02     *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:77        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:78        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:79        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:80N       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:81        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:86        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:88        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:89        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:96        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- -G- --- --- 
 DRB4*01:97        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- T-C TC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:98        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:99        *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:100       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
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 DRB4*01:104       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:105       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- -T- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:106       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:107       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:108N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:109       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:110       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:111       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:112       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:113N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- .-- --- --- --- 
 DRB4*01:114       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:115N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- -.. ..- --- --- C-- --- --- --- 
 DRB4*01:116       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:117       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:118       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:119       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:120       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:121N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- T-- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:122       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -C- --G --- --- --- C-- --- --- --- 
 DRB4*01:123       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:124       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:125       *** *** *** *** *** *** *** *** *|-- --- --- --- AA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:126       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:127       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:128N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:129       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:130       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:131       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- G-- --- C-- --- --- --- 
 DRB4*01:132       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:133       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- -G- 
 DRB4*01:134       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:135       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:136       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:137       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:138       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:139       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:140       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:141       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:142       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:143       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:144       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:145N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:146       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:147       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:148       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:149N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:150       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:151       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:152       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:153       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:154       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:155       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:156       --- --- --- --- --- --- --- -A- T|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:157       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -GG --G --- --- --- C-- --- --- --- 
 DRB4*01:158       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:159N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:160       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:161       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:162N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:163       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:164       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:165N      --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:166       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:167       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:168       --- --- --- -A- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:169       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:170       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:171       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:172       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:173       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:174       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:175       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:176       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:177       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:178       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:179       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:180       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:181       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:182       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:183       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:184       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:185       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:186       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:187       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:188       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:189       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:190       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:191       --- --- --C --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:192N      *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*01:193       *** *** *** *** *** *** *** *** *|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*02:01N       --- --- --- --- --- --- --- -A- -|-- --- --- --- GA- --- GC- --- -G- --G --- --- --- C-- --- --- --- 
 DRB4*03:01N       ... ... ... ... ... ... ... ... .|.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... 
                   
 cDNA              151
 AA codon          22
                   |
 DRB1*01:01:01:01  GAG CGG GTG CGG TTG CTG GAA AGA TGC ATC TAT AAC CAA GAG GAG TCC GTG CGC TTC GAC AGC GAC GTG GGG GAG 
 DRB4*01:01:01:01  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:01:03  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:01:04  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:01:05  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:01:06  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:02     --- --A --- T-- AAC --- ATC --- -A- --- --C --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:03     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:04     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:05     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:06     --A --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:07     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:08     --- A-A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:09     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -AT -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:10     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:11     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:12     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:13     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:01:14     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:02:01:01  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:02:01:02  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:01  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:02N --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:03  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:04  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:05  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:06  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:07  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:08  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:09  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:10  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:11  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:12  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:13N --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:14  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:15  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:16  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:17  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:18  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:19  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:20  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:21  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:22  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:23  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:24  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:25  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:26N --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:27  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:28N --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:29  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:01:30  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:01  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:02  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:03  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:04  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:05  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:06  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:07  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:08  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:09  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:02:10  --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:03     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:04     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --T C-- --- --- 
 DRB4*01:03:05     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:06     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:07     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:08     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:09     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:10     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:11     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:12     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:13     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -CC --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:14     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:15     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:16     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:17     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:18     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:19     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:20     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:21     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:22     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:23     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -CT --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:24     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:25     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:26     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:27     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:28     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:29     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- T-- --- --- 
 DRB4*01:03:30     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:31     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:32     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --A -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:33     --- --A --- T-- AAC --- ATC --- -A- --- --- --T --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:34     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:35     --- --A --- T-- AAC --C ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:36     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:03:37     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:04        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:05        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:06        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:07:01     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:07:02     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:08        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:09        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:10        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- AC- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:11        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:12        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:13        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:14N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:15        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:16N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:17        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:18        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:19        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:20        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:21        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:22        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:23        --- -AA --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:24        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:25        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:26        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:27        --- --A --- T-- AAC --- AGC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:28        --- --A --- T-- AAC --- ATC --- CA- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:29        --- --A --- T-- AAC --- ATC --- GA- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:30        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --C -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:31        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- CA- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:32        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- -A- -A- A-- --T --- C-- --- --- 
 DRB4*01:33        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- A-- 
 DRB4*01:34        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:35        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:36        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:37        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:38N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:39        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:40        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:41        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:42        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:43        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:44        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- T-- -A- A-- --T --- C-- --- --- 
 DRB4*01:45        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:46        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:47        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:48        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:49        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:50        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:51        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:52        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:53        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:54:01N    --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:54:02N    --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:55        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:56N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:57N       --- --A --- T-- AAC --- ATC --- -A- --- --G --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:58        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:59        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:60        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:61:01N    --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:61:02N    --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:62        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- --- --- --- --- --- --- --- 
 DRB4*01:63        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -G- A-- --T --- C-- --- --- 
 DRB4*01:64        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:65N       --- T-A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:66        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:67        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:68        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:69        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:70        --- --A --- T-- AAC --- TTC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:71N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:72:01     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:72:02     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:73        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- --- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:74        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:75        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:76:01     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:76:02     --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:77        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:78        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- -T- -A- A-- --T --- C-- --- --- 
 DRB4*01:79        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:80N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:81        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:82        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:83        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:84N       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:85        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:86        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:87        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:88        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --- --- --- --- --- 
 DRB4*01:89        --- --A --- T-- AAA --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:90        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:91        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:92        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:93        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- G-T --- C-- --- --- 
 DRB4*01:94        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:95        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:96        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:97        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:98        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:99        --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- -G- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:100       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:101       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:102       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:103       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:104       --- --A --- T-- AAC --- ATC --- -A- --- -T- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:105       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:106       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:107       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:108N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- -.T --- C-- --- --- 
 DRB4*01:109       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:110       A-- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:111       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:112       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -C- A-- --T --- C-- --- --- 
 DRB4*01:113N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:114       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:115N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:116       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- AG- --T --- C-- --- --- 
 DRB4*01:117       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- AA- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:118       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:119       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:120       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- A-- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:121N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:122       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:123       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:124       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:125       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:126       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:127       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:128N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:129       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- -T- -A- A-- --T --- C-- --- --- 
 DRB4*01:130       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:131       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:132       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:133       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:134       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:135       --- --A --- T-- AAC --- ATC --- -A- --G --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:136       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:137       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:138       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:139       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:140       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:141       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:142       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:143       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:144       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:145N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:146       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:147       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:148       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:149N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:150       --- --A -C- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:151       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:152       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:153       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:154       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:155       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:156       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:157       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:158       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:159N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:160       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:161       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:162N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:163       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:164       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:165N      --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:166       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:167       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:168       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:169       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:170       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:171       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:172       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:173       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:174       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:175       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:176       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:177       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:178       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:179       --- --A --- T-- AAC --- ATC --- -C- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:180       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:181       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:182       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:183       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- C-- --T --- C-- --- --- 
 DRB4*01:184       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:185       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:186       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:187       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:188       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:189       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:190       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:191       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:192N      --- --A --- T-- AAC --- ATC --- -AG --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*01:193       --- --A --- T-- AAC --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*02:01N       --- -.. ... ... ... --- ATC --- -A- --- --- --- --- --- --- -A- -C- --- -A- A-- --T --- C-- --- --- 
 DRB4*03:01N       ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... 
                   
 cDNA              226
 AA codon          47
                   |
 DRB1*01:01:01:01  TAC CGG GCG GTG ACG GAG CTG GGG CGG CCT GAT GCC GAG TAC TGG AAC AGC CAG AAG GAC CTC CT.G GAG CAG AGG 
 DRB4*01:01:01:01  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:01:03  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:01:04  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:01:05  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:01:06  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:02     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:03     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:04     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:05     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:06     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:07     --- -A- --A --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:08     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:09     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:10     --- -A- --- --C --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:11     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:12     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:13     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:01:14     --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:02:01:01  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:02:01:02  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:01  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:02N --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:03  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:04  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:05  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
 DRB4*01:03:01:06  --- -A- --- --- --- --- --- --- --- --- --C --T --- --- --- --- --- --- --- --- --- --.- --- -G- --- 
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 DRB4*03:01N       ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... .|-C C-A 
                   
 cDNA              376
 AA codon          97
                   |
 DRB1*01:01:01:01  CCT AAG GTG ACT GTG TAT CCT TCA AAG ACC CAG CCC CTG CAG CAC CAC AAC CTC CTG GTC TGC TCT GTG AGT GGT 
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 DRB4*01:189       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
 DRB4*01:190       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
 DRB4*01:191       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
 DRB4*01:192N      --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
 DRB4*01:193       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- T-- --- --- --- --- --- 
 DRB4*02:01N       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
 DRB4*03:01N       --- --- --- --- --- --C --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- --- 
                   
 cDNA              601
 AA codon          172
                   |
 DRB1*01:01:01:01  ACC TGC CAA GTG GAG CAC CCA AGT GTG ACG AGC CCT CTC ACA GTG GAA TGG A|GA GCA CGG TCT GAA TCT GCA CAG 
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 cDNA              676
 AA codon          197
                   |
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                   |
 DRB1*01:01:01:01  AGG AAT CAG AAA G|GA CAC TCT GGA CTT CAG CCA ACA G|GA TTC CTG AGC TGA         
 DRB4*01:01:01:01  --- --- --- --- -|-- --- --- --- --- --- --- --- -|-- C-- T-- --- ---         
 DRB4*01:01:01:03  --- --- --- --- -|-- --- --- --- --- --- --- --- -|-- C-- T-- --- ---         
 DRB4*01:01:01:04  --- --- --- --- -|-- --- --- --- --- --- --- --- -|-- C-- T-- --- ---         
 DRB4*01:01:01:05  --- --- --- --- -|-- --- --- --- --- --- --- --- -|-- C-- T-- --- ---         
 DRB4*01:01:01:06  --- --- --- --- -|-- --- --- --- --- --- --- --- -|-- C-- T-- --- ---         
 DRB4*01:01:02     *** *** *** *** *|** *** *** *** *** *** *** *** *|** *** *** *** ***         
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