The post-translational modifications used in the UniProt knowledgebase (Swiss-Prot and TrEMBL). The definition of the post-translational modifications usage as well as other information is provided in the following format CC-BY-4.0 UniProt Post-Translational Modification help@uniprot.org PyOBO v0.12.13-dev-207b46e9 on 2025-10-31T15:17:16.572734 by https://github.com/biopragmatics/pyobo/blob/main/src/pyobo/sources/uniprot/uniprot_ptm.py 2025_04 https://ftp.uniprot.org/pub/databases/uniprot/current_release/knowledgebase/complete/docs/ptmlist https://www.uniprot.org/uniprot-logo.img.0df091.svg (E)-2,3-didehydrotyrosine (Z)-2,3-didehydrotyrosine 1'-histidyl-3'-tyrosine (His-Tyr) 1-thioglycine 2'-(S-cysteinyl)-histidine (Cys-His) 2,3-didehydroalanine (Ser) 2,3-didehydrobutyrine 2,3-didehydrotyrosine 2',4',5'-topaquinone 2-cysteinyl-D-allo-threonine (Cys-Thr) 2-cysteinyl-D-phenylalanine (Cys-Phe) 2-cysteinyl-L-phenylalanine (Cys-Phe) 2-iminomethyl-5-imidazolinone (Gln-Gly) 2-iminomethyl-5-imidazolinone (Glu-Gly) 2-iminomethyl-5-imidazolinone (Met-Gly) 2-methylglutamine 2-oxobutanoic acid 2-tetrahydro-2-pyridyl-5-imidazolinone (Lys-Gly) 3'-(S-cysteinyl)-tyrosine (Cys-Tyr) 3-(S-cysteinyl)-tyrosine (Cys-Tyr) 3'-(S-cysteinyl)-tyrosine (Tyr-Cys) 3,4-dihydroxyarginine 3',4'-dihydroxyphenylalanine 3,4-dihydroxyproline 3-cysteinyl-aspartic acid (Cys-Asp) 3'-geranyl-2',N2-cyclotryptophan 3'-histidyl-3-tyrosine (His-Tyr) 3-hydroxyasparagine 3-hydroxyaspartate 3-hydroxyproline 3-hydroxytryptophan 3-methylthioaspartic acid 3-oxoalanine (Cys) 3-oxoalanine (Ser) 3-phenyllactic acid 4,5-dihydroxylysine N,N-(cysteine-1,S-diyl)serine (Cys-Ser) 4-aspartylphosphate 4-carboxyglutamate 4-cysteinyl-glutamic acid (Cys-Glu) 4'-cysteinyl-tryptophylquinone (Cys-Trp) 4-hydroxyarginine 4-hydroxyproline 5-hydroxylysine 5-imidazolinone (Ala-Gly) 5-imidazolinone (Asn-Gly) 5-imidazolinone (Cys-Gly) 5-imidazolinone (Lys-Gly) 5-imidazolinone (Ser-Gly) 5-methylarginine 6'-bromotryptophan 6'-chlorotryptophan ADP-ribosylarginine ADP-ribosylasparagine ADP-ribosylcysteine ADP-ribosylserine Alanine amide Alanine derivative Allysine Arginine amide Arginine derivative Asparagine amide Aspartic acid 1-amide Aspartyl aldehyde Asymmetric dimethylarginine Beta-methyllanthionine (Cys-Thr) Beta-methyllanthionine (Thr-Cys) Beta-methyllanthionine sulfoxide (Thr-Cys) Blocked amino end (Ala) Blocked amino end (Arg) Blocked amino end (Asn) Blocked amino end (Asp) Blocked amino end (Asx) Blocked amino end (Cys) Blocked amino end (Gln) Blocked amino end (Glu) Blocked amino end (Gly) Blocked amino end (Ile) Blocked amino end (Leu) Blocked amino end (Met) Blocked amino end (Pro) Blocked amino end (Ser) Blocked amino end (Thr) Blocked amino end (Val) Blocked amino end (Xaa) Blocked carboxyl end (Arg) Blocked carboxyl end (His) Bromohistidine Cholesterol glycine ester Cis-14-hydroxy-10,13-dioxo-7-heptadecenoic acid aspartate ester Citrulline Cyclopeptide (Ala-Arg) Cyclopeptide (Arg-Cys) (interchain with C-...) Cyclopeptide (Asn-Gly) Cyclopeptide (Asp-Asn) Cyclopeptide (Cys-Arg) (interchain with R-...) Cyclopeptide (Gly-Asn) Cyclopeptide (Gly-Asp) Cyclopeptide (Lys-Asp) Cyclopeptide (Ser-Asn) Cysteine amide Cysteine derivative Cysteine methyl disulfide Cysteine methyl ester Cysteine persulfide Cysteine sulfenic acid (-SOH) Cysteine sulfinic acid (-SO2H) Cysteinyl-selenocysteine (Cys-Sec) Cysteinyl-selenocysteine (Sec-Cys) D-4-hydroxyvaline D-alanine (Ala) D-alanine (Ser) D-allo-isoleucine D-asparagine Deamidated asparagine Deamidated glutamine Diphthamide D-leucine D-methionine D-phenylalanine D-tryptophan D-valine FMN phosphoryl serine FMN phosphoryl threonine Glutamate methyl ester (Gln) Glutamate methyl ester (Glu) Glutamic acid 1-amide Glutamine amide Glutamine derivative Glycine amide Glycine radical Glycyl lysine isopeptide (Gly-Lys) (interchain with K-...) Glycyl lysine isopeptide (Lys-Gly) (interchain with G-...) GPI-anchor amidated alanine GPI-anchor amidated asparagine GPI-anchor amidated aspartate GPI-anchor amidated carboxyl end GPI-anchor amidated cysteine GPI-anchor amidated glycine GPI-anchor amidated serine GPI-anchor amidated threonine GPI-like-anchor amidated alanine GPI-like-anchor amidated asparagine GPI-like-anchor amidated glycine GPI-like-anchor amidated serine Histidine amide Hydroxyproline Hypusine Isoaspartyl cysteine isopeptide (Cys-Asn) Isoaspartyl glycine isopeptide (Gly-Asn) Isoaspartyl lysine isopeptide (Asn-Lys) (interchain with K-...) Isoaspartyl lysine isopeptide (Lys-Asn) (interchain with N-...) Isodityrosine (Tyr-Tyr) Isoglutamyl cysteine thioester (Cys-Gln) Isoglutamyl glycine isopeptide (Gly-Glu) Isoglutamyl lysine isopeptide (Gln-Lys) (interchain with K-...) Isoglutamyl lysine isopeptide (Lys-Gln) Isoglutamyl lysine isopeptide (Lys-Gln) (interchain with Q-...) Isoleucine amide Isoleucine derivative D-lactate Lanthionine (Cys-Ser) Lanthionine (Ser-Cys) Leucine amide Leucine methyl ester Lysine amide Lysine derivative Lysine methyl ester Lysine tyrosylquinone (Lys-Tyr) Lysinoalanine (Ser-Lys) Methionine amide Methionine derivative Methionine sulfone Methylhistidine N,N,N-trimethylalanine N,N-dimethylalanine N,N-dimethylproline N2-acetylarginine N2-succinyltryptophan N4,N4-dimethylasparagine N4-methylasparagine N5-methylarginine N5-methylglutamine N6,N6,N6-trimethyl-5-hydroxylysine N6,N6,N6-trimethyllysine N6,N6-dimethyllysine N6-1-carboxyethyl lysine N6-acetyllysine N6-carboxylysine N6-formyllysine N6-methylated lysine N6-methyllysine N6-murein peptidoglycan lysine N6-myristoyl lysine N6-palmitoyl lysine N6-poly(methylaminopropyl)lysine N-acetylalanine N-acetylaspartate N-acetylcysteine N-acetylglutamate N-acetylglycine N-acetylisoleucine N-acetylmethionine N-acetylproline N-acetylserine N-acetylthreonine N-acetyltyrosine N-acetylvaline N-formylglycine N-formylmethionine Nitrated tyrosine N-methylalanine N-methylisoleucine N-methylleucine N-methylmethionine N-methylphenylalanine N-methylproline N-methyltyrosine N-myristoyl glycine N-palmitoyl cysteine N-palmitoyl glycine N-pyruvate 2-iminyl-cysteine N-pyruvate 2-iminyl-valine O-(5'-phospho-DNA)-serine O-(5'-phospho-RNA)-serine O-(5'-phospho-DNA)-tyrosine O-(5'-phospho-RNA)-tyrosine O-(sn-1-glycerophosphoryl)serine O-8alpha-FAD tyrosine O-acetylserine O-acetylthreonine O-decanoyl serine O-decanoyl threonine Omega-hydroxyceramide glutamate ester Omega-N-methylarginine Omega-N-methylated arginine O-octanoyl serine O-octanoyl threonine O-palmitoyl serine O-palmitoyl threonine Pentaglycyl murein peptidoglycan amidated alanine Pentaglycyl murein peptidoglycan amidated threonine Peptide (Met-Gly) (interchain with G-...) Phenylalanine amide Phosphatidylethanolamine amidated glycine Phosphoarginine Phosphocysteine Phosphohistidine Phosphoserine Phosphothreonine Phosphotyrosine PolyADP-ribosyl glutamic acid Proline amide Pros-8alpha-FAD histidine Pros-methylhistidine Pros-phosphohistidine Pyrrolidone carboxylic acid Pyrrolidone carboxylic acid (Glu) Pyrroloquinoline quinone (Glu-Tyr) Pyruvic acid (Cys) Pyruvic acid (Ser) S-(2-aminovinyl)-3-methyl-D-cysteine (Thr-Cys) S-(2-aminovinyl)-D-cysteine (Ser-Cys) S-12-hydroxyfarnesyl cysteine S-4a-FMN cysteine S-6-FMN cysteine S-8alpha-FAD cysteine S-archaeol cysteine S-diacylglycerol cysteine Serine amide Serine microcin E492 siderophore ester S-farnesyl cysteine S-geranylgeranyl cysteine S-methylcysteine S-nitrosocysteine S-palmitoyl cysteine S-selanylcysteine S-stearoyl cysteine Sulfoserine Sulfothreonine Sulfotyrosine Symmetric dimethylarginine Tele-8alpha-FAD histidine Tele-8alpha-FMN histidine Tele-methylhistidine Threonine amide Thyroxine Triiodothyronine Tryptophan amide Tryptophan derivative Tryptophan tryptophylquinone (Trp-Trp) Tryptophylquinone Tyrosine amide Valine amide 2'-methylsulfonyltryptophan (3R,4S)-3,4-dihydroxyproline Cyclopeptide (Ala-Pro) D-serine (Ser) D-serine (Cys) D-threonine S-glutathionyl cysteine (2-aminosuccinimidyl)acetic acid (Asp-Gly) Beta-decarboxylated aspartate Cyclopeptide (Ala-Ile) Cyclopeptide (Pro-Met) Cyclopeptide (Pro-Tyr) Cyclopeptide (Ser-Gly) Cyclopeptide (Leu-Trp) 5'-tyrosyl-5'-aminotyrosine (Tyr-Tyr) (interchain with Y-...) Aminomalonic acid (Ser) GPI-like-anchor amidated aspartate Lysine tyrosylquinone (Tyr-Lys) S-cysteinyl 3-(oxidosulfanyl)alanine (Cys-Cys) Tele-phosphohistidine Threonyl lysine isopeptide (Lys-Thr) (interchain with T-...) Threonyl lysine isopeptide (Thr-Lys) (interchain with K-...) Tryptophyl-tyrosyl-methioninium (Trp-Tyr) (with M-...) Tryptophyl-tyrosyl-methioninium (Tyr-Met) (with W-...) Isoaspartyl lysine isopeptide (Lys-Asn) N-D-glucuronoyl glycine O-AMP-tyrosine O-UMP-tyrosine Aspartate 1-(chondroitin 4-sulfate)-ester Aspartic acid 1-[(3-aminopropyl)(5'-adenosyl)phosphono]amide (3R,4R)-4,5-dihydroxyisoleucine (3R,4S)-4-hydroxyisoleucine 2'-cysteinyl-6'-hydroxytryptophan sulfoxide (Trp-Cys) Cyclopeptide (Ile-Pro) (4R)-4,5-dihydroxyleucine Dimethylated arginine (3S,4R)-3,4-dihydroxyisoleucine 3-hydroxyphenylalanine 3-hydroxyvaline (Val) 5-amino-piperideine-2,5-dicarboxylic acid (Ser-Cys) (with S-...) 5-amino-piperideine-2,5-dicarboxylic acid (Ser-Ser) (with C-...) 5-methylthiazole-4-carboxylic acid (Asn-Cys) Cyclopeptide (Leu-Leu) Decarboxylated threonine N6-(ADP-ribosyl)lysine O-methylthreonine Pyridine-2,5-dicarboxylic acid (Ser-Cys) (with S-...) Pyridine-2,5-dicarboxylic acid (Ser-Ser) (with C-...) Thiazole-4-carboxylic acid (Asn-Cys) Thiazole-4-carboxylic acid (Cys-Cys) Thiazole-4-carboxylic acid (Ile-Cys) Thiazole-4-carboxylic acid (Pro-Cys) Thiazole-4-carboxylic acid (Ser-Cys) Thiazole-4-carboxylic acid (Thr-Cys) Thiazole-4-carboxylic acid (Val-Cys) Thiazoline-4-carboxylic acid (Phe-Cys) (3R,4R)-3,4-dihydroxyproline (3R)-3-hydroxyasparagine (3S)-3-hydroxyasparagine (3R)-3-hydroxyaspartate 4,5,5'-trihydroxyleucine 5-(methoxymethyl)thiazole-4-carboxylic acid (Val-Cys) N-carbamoylalanine Thiazole-4-carboxylic acid (Phe-Cys) Oxazole-4-carboxylic acid (Cys-Ser) Oxazole-4-carboxylic acid (Gly-Ser) Thiazole-4-carboxylic acid (Gly-Cys) 1-amino-2-propanone Cyclo[(prolylserin)-O-yl] cysteinate Oxazoline-4-carboxylic acid (Cys-Ser) N6-biotinyllysine N6-lipoyllysine 5-methyloxazole-4-carboxylic acid (Ser-Thr) N6-(pyridoxal phosphate)lysine N6-(retinylidene)lysine O-(phosphoribosyl dephospho-coenzyme A)serine 3-(O4'-tyrosyl)-valine (Val-Tyr) O-(pantetheine 4'-phosphoryl)serine (4S)-thiazoline-4-carboxylic acid (Thr-Cys) O-AMP-threonine 5-glutamyl polyglycine 5-glutamyl polyglutamate Isoglutamyl lysine isopeptide (Glu-Lys) (interchain with K-...) Isoglutamyl lysine isopeptide (Lys-Glu) (interchain with E-...) O-(2-aminoethylphosphoryl)serine O-(2-cholinephosphoryl)serine S-Lysyl-methionine sulfilimine (Met-Lys) (interchain with K-...) S-Lysyl-methionine sulfilimine (Lys-Met) (interchain with M-...) 5-glutamyl glycerylphosphorylethanolamine 5-glutamyl 2-aminoadipic acid 5-glutamyl N2-lysine Diiodotyrosine Glycyl adenylate Glycyl cysteine dithioester (Gly-Cys) (interchain with C-...) Iodotyrosine Threonine methyl ester Glycyl cysteine dithioester (Cys-Gly) (interchain with G-...) S-(4-hydroxycinnamyl)cysteine S-cysteinyl cysteine Tele-(1,2,3-trihydroxypropan-2-yl)histidine Trithiocysteine (Cys-Cys) Alanine isoaspartyl cyclopeptide (Ala-Asn) N-[(12R)-12-hydroxymyristoyl]cysteine N-(12-oxomyristoyl)cysteine S-(dipyrrolylmethanemethyl)cysteine Glycyl serine ester (Ser-Gly) (interchain with G-...) Glycyl threonine ester (Thr-Gly) (interchain with G-...) 2-(S-cysteinyl)pyruvic acid O-phosphothioketal Glycyl serine ester (Gly-Ser) (interchain with S-...) Glycyl threonine ester (Gly-Thr) (interchain with T-...) 7'-hydroxytryptophan S-(coelenterazin-3a-yl)cysteine Glycyl cysteine thioester (Cys-Gly) (interchain with G-...) N,N,N-trimethylserine N,N-dimethylserine N-methylserine CysO-cysteine adduct 3'-nitrotyrosine N,N-dimethylleucine Cyclopeptide (His-Asn) Cyclopeptide (His-Asp) N6-succinyllysine Lysino-D-alanine (Lys) (Z)-2,3-didehydrobutyrine (E)-2,3-didehydrobutyrine L-allo-isoleucine S-(2-aminovinyl)-L-cysteine (Cys-Cys) 5'-chlorotryptophan 1-(tryptophan-3-yl)-tryptophan (Trp-Trp) (interchain with W-...) S-(2-aminovinyl)-D-cysteine (Cys-Cys) S-(15-deoxy-Delta12,14-prostaglandin J2-9-yl)cysteine 2-(3-methylbutanoyl)-5-hydroxyoxazole-4-carbothionic acid (Leu-Cys) Proline 5-hydroxy-oxazole-4-carbothionic acid (Pro-Cys) (2-aminosuccinimidyl)acetic acid (Asn-Gly) N,N-(cysteine-1,S-diyl)phenylalanine (Cys-Phe) S-bacillithiol cysteine disulfide 4-hydroxyglutamate 3-hydroxypyridine-2,5-dicarboxylic acid (Ser-Cys) (with S-...) 3-hydroxypyridine-2,5-dicarboxylic acid (Ser-Ser) (with C-...) Thiazole-4-carboxylic acid (Glu-Cys) 2-(4-guanidinobutanoyl)-5-hydroxyimidazole-4-carbothionic acid (Arg-Cys) Threonine 5-hydroxy-oxazole-4-carbonthionic acid (Thr-Cys) N2,N2-dimethylarginine Thiazole-4-carboxylic acid (Arg-Cys) 5-methyloxazole-4-carboxylic acid (Cys-Thr) 5-methyloxazole-4-carboxylic acid (Thr-Thr) Oxazole-4-carboxylic acid (Ile-Ser) Oxazole-4-carboxylic acid (Ser-Ser) 5-methyloxazoline-4-carboxylic acid (Ser-Thr) 5-hydroxy-3-methylproline (Ile) N6-malonyllysine 2,3-didehydroalanine (Cys) Methionine sulfoxide (5R)-5-hydroxylysine (5S)-5-hydroxylysine (3S)-3-hydroxyaspartate N6-(3,6-diaminohexanoyl)-5-hydroxylysine N6-crotonyllysine (3R)-3-hydroxyarginine (3S)-3-hydroxyhistidine 5-glutamyl N2-ornithine 5-glutamyl glutamate Methionine (R)-sulfoxide Methionine (S)-sulfoxide N-methylglycine N,N-dimethylglycine N,N,N-trimethylglycine Isoaspartyl lysine isopeptide (Lys-Asp) N6-glutaryllysine 5-glutamyl histamine Isoaspartyl glycine isopeptide (Asn-Gly) Isoaspartyl glycine isopeptide (Asp-Gly) (4R)-5-hydroxyleucine (4R)-5-oxoleucine O-palmitoleoyl serine PolyADP-ribosyl aspartic acid 2-(S-cysteinyl)-methionine (Cys-Met) Cyclopeptide (Cys-Ile) ADP-ribosyl aspartic acid Hydroxyarginine N6-(beta-hydroxybutyryl)lysine O-UMP-histidine O-UMP-serine O-UMP-threonine N,N,N-trimethylmethionine C-linked (Man) hydroxytryptophan C-linked (Man) tryptophan N-linked (DATDGlc) asparagine N-linked (Glc) (glycation) histidine N-linked (Glc) (glycation) isoleucine N-linked (Glc) (glycation) lysine N-linked (Glc) (glycation) valine N-linked (Lac) (glycation) lysine N-linked (GalNAc) asparagine N-linked (GalNAc...) asparagine N-linked (GalNAc...) (glycosaminoglycan) asparagine N-linked (Glc) arginine N-linked (Glc...) arginine N-linked (Glc) asparagine N-linked (Glc...) asparagine N-beta-linked (GlcNAc) arginine N-linked (GlcNAc...) arginine N-linked (GlcNAc...) (complex) arginine N-linked (GlcNAc...) (hybrid) arginine N-linked (GlcNAc...) (high mannose) arginine N-linked (GlcNAc...) (paucimannose) arginine N-linked (GlcNAc...) (keratan sulfate) arginine N-linked (GlcNAc...) (polylactosaminoglycan) arginine N-linked (GlcNAc) asparagine N-linked (GlcNAc...) asparagine N-linked (GlcNAc...) (complex) asparagine N-linked (GlcNAc...) (hybrid) asparagine N-linked (GlcNAc...) (high mannose) asparagine N-linked (GlcNAc...) (paucimannose) asparagine N-linked (GlcNAc...) (keratan sulfate) asparagine N-linked (GlcNAc...) (polylactosaminoglycan) asparagine N-linked (Man) tryptophan N-linked (Hex) arginine N-linked (Hex...) arginine N-linked (HexNAc) arginine N-linked (HexNAc...) arginine N-linked (Hex) asparagine N-linked (Hex...) asparagine N-linked (HexNAc) asparagine N-linked (HexNAc...) asparagine N-linked (Hex) tryptophan O-linked (Ara) hydroxyproline O-linked (Ara...) hydroxyproline O-linked (DADDGlc) serine O-linked (DATDGlc) serine O-linked (GATDGlc) serine O-linked (Fuc) serine O-linked (Fuc...) serine O-linked (Fuc) threonine O-linked (Fuc...) threonine O-linked (FucNAc) serine O-linked (FucNAc...) serine O-linked (Gal) hydroxylysine O-linked (Gal...) hydroxylysine O-linked (Gal) hydroxyproline O-linked (Gal...) hydroxyproline O-linked (Gal) serine O-linked (Gal...) serine O-linked (Gal) threonine O-linked (Gal...) threonine O-linked (GalNAc) serine O-linked (GalNAc...) serine O-linked (GalNAc...) (keratan sulfate) serine O-linked (GalNAc) threonine O-linked (GalNAc...) threonine O-linked (GalNAc...) (keratan sulfate) threonine O-linked (GalNAc) tyrosine O-linked (GalNAc...) tyrosine O-linked (Glc) hydroxylysine O-linked (Glc) serine O-linked (Glc...) serine O-linked (Glc) tyrosine O-linked (Glc...) tyrosine O-linked (GlcA) serine O-linked (GlcNAc) hydroxyproline O-linked (GlcNAc...) hydroxyproline O-linked (GlcNAc) serine O-linked (GlcNAc...) serine O-linked (GlcNAc) threonine O-linked (GlcNAc...) threonine O-linked (GlcNAc) tyrosine O-linked (GlcNAc...) tyrosine O-linked (GlcNAc1P) serine O-linked (GlcNAc6P) serine O-linked (Man) serine O-linked (Man...) serine O-linked (Man...) (keratan sulfate) serine O-linked (Man) threonine O-linked (Man...) threonine O-linked (Man...) (keratan sulfate) threonine O-linked (Man1P) serine O-linked (Man1P...) serine O-linked (Man6P) threonine O-linked (Man6P...) threonine O-linked (Xyl) serine O-linked (Xyl...) serine O-linked (Xyl...) (chondroitin sulfate) serine O-linked (Xyl...) (dermatan sulfate) serine O-linked (Xyl...) (heparan sulfate) serine O-linked (Xyl...) (glycosaminoglycan) serine O-linked (Xyl...) (keratan sulfate) threonine O-linked (Xyl...) (glycosaminoglycan) threonine O-linked (Hex) hydroxylysine O-linked (Hex...) hydroxylysine O-linked (Hex) hydroxyproline O-linked (Hex...) hydroxyproline O-linked (Hex) serine O-linked (Hex...) serine O-linked (Hex) threonine O-linked (Hex...) threonine O-linked (Hex) tyrosine O-linked (Hex...) tyrosine O-linked (HexNAc) hydroxyproline O-linked (HexNAc...) hydroxyproline O-linked (HexNAc) serine O-linked (HexNAc...) serine O-linked (HexNAc) threonine O-linked (HexNAc...) threonine O-linked (HexNAc) tyrosine O-linked (HexNAc...) tyrosine S-linked (Gal) cysteine S-linked (Gal...) cysteine S-linked (Glc) cysteine S-linked (Glc...) cysteine S-linked (GlcNAc) cysteine S-linked (GlcNAc...) cysteine S-linked (Hex) cysteine S-linked (Hex...) cysteine S-linked (HexNAc) cysteine S-linked (HexNAc...) cysteine Cysteine sulfonic acid (-SO3H) Cyclopeptide (Glu-Asn) S-methylmethionine N6-butyryllysine N6-(2-hydroxyisobutyryl)lysine N6-poly(beta-hydroxybutyryl)lysine O3-poly(beta-hydroxybutyryl)serine S-poly(beta-hydroxybutyryl)cysteine N6-propionyllysine Cyclopeptide (Gly-Arg) Cyclopeptide (Ser-Lys) S-palmitoleoyl cysteine ADP-ribosyl glutamic acid 2,3-didehydroalanine (Tyr) Glycyl cysteine thioester (Gly-Cys) (interchain with C-...) S-carbamoylcysteine S-cyanocysteine O-AMP-serine Cyclopeptide (Cys-Pro) Cyclopeptide (Gly-Pro) Cyclopeptide (His-Pro) Cyclopeptide (Leu-Pro) Cyclopeptide (Phe-Pro) Cyclopeptide (Tyr-Pro) Cyclopeptide (Ser-Pro) Cyclopeptide (Trp-Pro) Cyclopeptide (Val-Pro) Cyclopeptide (Met-Pro) Pyruvic acid (Tyr) (3S)-3-hydroxylysine 4-hydroxylysine (2S)-4-hydroxyleucine (4S)-4,5-dihydroxyleucine 3',4',5'-trihydroxyphenylalanine 2-hydroxyproline Murein peptidoglycan amidated serine ADP-ribosyltyrosine ADP-ribosylglycine ADP-ribosyldiphthamide (Z)-2,3-didehydroaspartate S-(2-succinyl)cysteine N6-carbamoyllysine S-(2,3-dicarboxypropyl)cysteine S-cGMP-cysteine N5-[4-(S-L-cysteinyl)-5-methyl-1H-imidazol-2-yl]-L-ornithine (Arg-Cys) (interchain with C-...) N5-[4-(S-L-cysteinyl)-5-methyl-1H-imidazol-2-yl]-L-ornithine (Cys-Arg) (interchain with R-...) N-linked (Glc) (glycation) arginine 6-(S-cysteinyl)-8alpha-(pros-histidyl)-FAD (His-Cys) N6-lactoyllysine Thiazolidine linkage to a ring-opened DNA abasic site Deoxyhypusine 3'-prenyl-2',N2-cyclotryptophan Isoaspartyl glycine isopeptide (Gly-Asp) 2-oxo-5,5-dimethylhexanoate 3-methylisoleucine 3-methylvaline 3-methyl-D-valine N4-methyl-D-asparagine 3-hydroxyvaline (Thr) (3S)-3-methylglutamine 3-hydroxy-D-valine (3R)-N4-methyl-3-hydroxy-D-asparagine 3,3-dimethylmethionine Diphosphoserine Diphosphothreonine O-linked (Glc) threonine O-alpha-linked (GlcNAc) threonine O-di-AMP-tyrosine O-tri-AMP-tyrosine 5-glutamyl dopamine 5-glutamyl noradrenaline 5-glutamyl serotonin N-alpha-linked (Rha) arginine 5-glutamyl glycine ADP-alpha-D-ribosylarginine Phosphatidylserine amidated glycine 4-(1-hydroxyethyl)-7-isoleucino-2-(threonin-O3-ylcarbonyl)-7,8-dihydroquinolin-8-ol (Ile-Thr) O-hexanoyl serine 3'-farnesyl-2',N2-cyclotryptophan Cyclopeptide (Ile-Lys) ADP-riboxanated arginine 3'-bromotyrosine N6-stearoyl lysine 2-(cystein-S-ylcarbonyl)-3-methyl-4-(glutam-5-yloxy)methylindole (Glu-Cys) Dityrosine (Tyr-Tyr) (interchain with Y-...) ADP-ribosylhistidine ADP-ribosylthreonine N6-decanoyllysine O-alpha-linked (D-glycero-D-manno-heptose) serine O-alpha-linked (glycero-D-manno-heptose) serine O-alpha-linked (glycero-D-manno-heptose) threonine N-methylvaline N-methylthreonine N-methylaspartate N-methylhistidine Sulfocysteine L-cysteine coenzyme A disulfide Aspartate methyl ester N6-acetyl-N6-methyllysine GPI-anchor amidated isoleucine 4-hydroxynonenal-conjugated cysteine 4-hydroxynonenal-conjugated histidine 4-hydroxynonenal-conjugated lysine Methylcyclopropylglycine 3-(L-phenylalan-2'-yl)-L-valine (Phe-Val) Glyoxylic acid (Cys) N6-benzoyllysine N6-isonicotinyllysine N6-methacryllysine