István Mikó (imiko) https://orcid.org/0000-0003-2938-9075 Jennifer C. Girón (jgiron) https://orcid.org/0000-0002-0851-6883 Luis A. González Montaña (lagonzalezmo) https://orcid.org/0000-0002-9136-9932 Sergei Tarasov (starasov) https://orcid.org/0000-0001-5237-2330 The ontology for the Anatomy of the Insect SkeletoMuscular system (AISM) contains terms used to describe the cuticle - as a single anatomical structure - and the skeletal muscle system, to be used in insect biodiversity research. Ontology for the Anatomy of the Insect SkeletoMuscular system 2021-08-01 This annotation property defines a "sensu", which is the combination of a bibliographic reference, a label and a class (represented by verbatim and logical definitions). Given this model, independent observations can be inferred across to compute synonymy (multiple labels attached to one), homonymy (one label attached to multiple classes). Yoder MJ, Mikó I, Seltmann KC, Bertone MA, Deans AR (2010) A Gross Anatomy Ontology for Hymenoptera. PLOS ONE 5: e15991. https://doi.org/10.1371/journal.pone.0015991.g004 This annotation property is on the label of a class in owl. sensu This annotation property defines a "sensu", which is the combination of a bibliographic reference, a label and a class (represented by verbatim and logical definitions). Given this model, independent observations can be inferred across to compute synonymy (multiple labels attached to one), homonymy (one label attached to multiple classes). Yoder MJ, Mikó I, Seltmann KC, Bertone MA, Deans AR (2010) A Gross Anatomy Ontology for Hymenoptera. PLOS ONE 5: e15991. https://doi.org/10.1371/journal.pone.0015991.g004 This annotation property is on the label of a class in owl. imiko https://orcid.org/0000-0003-2938-9075 2020-11-17 definition The official definition, explaining the meaning of a class or property. Shall be Aristotelian, formalized and normalized. Can be augmented with colloquial definitions. 2012-04-05: Barry Smith The official OBI definition, explaining the meaning of a class or property: 'Shall be Aristotelian, formalized and normalized. Can be augmented with colloquial definitions' is terrible. Can you fix to something like: A statement of necessary and sufficient conditions explaining the meaning of an expression referring to a class or property. Alan Ruttenberg Your proposed definition is a reasonable candidate, except that it is very common that necessary and sufficient conditions are not given. Mostly they are necessary, occasionally they are necessary and sufficient or just sufficient. Often they use terms that are not themselves defined and so they effectively can't be evaluated by those criteria. On the specifics of the proposed definition: We don't have definitions of 'meaning' or 'expression' or 'property'. For 'reference' in the intended sense I think we use the term 'denotation'. For 'expression', I think we you mean symbol, or identifier. For 'meaning' it differs for class and property. For class we want documentation that let's the intended reader determine whether an entity is instance of the class, or not. For property we want documentation that let's the intended reader determine, given a pair of potential relata, whether the assertion that the relation holds is true. The 'intended reader' part suggests that we also specify who, we expect, would be able to understand the definition, and also generalizes over human and computer reader to include textual and logical definition. Personally, I am more comfortable weakening definition to documentation, with instructions as to what is desirable. We also have the outstanding issue of how to aim different definitions to different audiences. A clinical audience reading chebi wants a different sort of definition documentation/definition from a chemistry trained audience, and similarly there is a need for a definition that is adequate for an ontologist to work with. PERSON:Daniel Schober GROUP:OBI:<http://purl.obolibrary.org/obo/obi> definition definition IAO:0000116 uberon editor_note true editor_note editor note editor note curator note An administrative note of use for a curator but of no use for a user PERSON:Alan Ruttenberg IAO:0000232 uberon curator_notes true curator_notes curator note curator note curator notes If R <- P o Q is a defining property chain axiom, then it also holds that R -> P o Q. Note that this cannot be expressed directly in OWL is a defining property chain axiom If R <- P o Q is a defining property chain axiom, then (1) R -> P o Q holds and (2) Q is either reflexive or locally reflexive. A corollary of this is that P SubPropertyOf R. is a defining property chain axiom where second argument is reflexive uberon dc-contributor true dc-contributor contributor uberon dc-description true dc-description description uberon dc-title true dc-title title uberon dcterms-license true dcterms-license license subset_property has_exact_synonym X lateral to y if x is further from the midsagittal plane than y. lateral_to X lateral to y if x is further from the midsagittal plane than y. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 lateral_to imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 X medial to y if x is closer to the midsagittal plane than y. medial_to X medial to y if x is closer to the midsagittal plane than y. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 medial_to imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 x encircles y if both x and y are regions of the 2D structure z, or 2D cross-sections of a 3D object z, and the boundary of y on z is exclusively continuous with x. encircles x encircles y if both x and y are regions of the 2D structure z, or 2D cross-sections of a 3D object z, and the boundary of y on z is exclusively continuous with x. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 encircles imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 y is encircled by x if both x and y are regions of the 2D structure z, or 2D cross-sections of a 3D object z, and the boundary of y on z is exclusively continuous with y. encircled by y is encircled by x if both x and y are regions of the 2D structure z, or 2D cross-sections of a 3D object z, and the boundary of y on z is exclusively continuous with y. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 encircled by imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 sclerite A encircles_via_conjunctiva sclerite B if A is encircled_via_conjuntiva X that encircles B encircles via conjunctiva sclerite A encircles_via_conjunctiva sclerite B if A is encircled_via_conjuntiva X that encircles B starasov https://orcid.org/0000-0001-5237-2330 2021-05-31 sclerite B is encircled_via_conjunctiva if sclerite A encircles_via_conjuntiva X sclerite B encircled via conjunctiva sclerite B is encircled_via_conjunctiva if sclerite A encircles_via_conjuntiva X sclerite B jgiron https://orcid.org/0000-0002-0851-6883 2021-06-14 is part of my brain is part of my body (continuant parthood, two material entities) my stomach cavity is part of my stomach (continuant parthood, immaterial entity is part of material entity) this day is part of this year (occurrent parthood) a core relation that holds between a part and its whole Everything is part of itself. Any part of any part of a thing is itself part of that thing. Two distinct things cannot be part of each other. Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See https://code.google.com/p/obo-relations/wiki/ROAndTime Parthood requires the part and the whole to have compatible classes: only an occurrent can be part of an occurrent; only a process can be part of a process; only a continuant can be part of a continuant; only an independent continuant can be part of an independent continuant; only an immaterial entity can be part of an immaterial entity; only a specifically dependent continuant can be part of a specifically dependent continuant; only a generically dependent continuant can be part of a generically dependent continuant. (This list is not exhaustive.) A continuant cannot be part of an occurrent: use 'participates in'. An occurrent cannot be part of a continuant: use 'has participant'. A material entity cannot be part of an immaterial entity: use 'has location'. A specifically dependent continuant cannot be part of an independent continuant: use 'inheres in'. An independent continuant cannot be part of a specifically dependent continuant: use 'bearer of'. part_of BFO:0000050 external protein quality spatial uberon part_of part_of part of part of part_of http://www.obofoundry.org/ro/#OBO_REL:part_of has part my body has part my brain (continuant parthood, two material entities) my stomach has part my stomach cavity (continuant parthood, material entity has part immaterial entity) this year has part this day (occurrent parthood) Q1 has_part Q2 if and only if: every instance of Q1 is a quality_of an entity that has_quality some Q2. a core relation that holds between a whole and its part Everything has itself as a part. Any part of any part of a thing is itself part of that thing. Two distinct things cannot have each other as a part. Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See https://code.google.com/p/obo-relations/wiki/ROAndTime Parthood requires the part and the whole to have compatible classes: only an occurrent have an occurrent as part; only a process can have a process as part; only a continuant can have a continuant as part; only an independent continuant can have an independent continuant as part; only a specifically dependent continuant can have a specifically dependent continuant as part; only a generically dependent continuant can have a generically dependent continuant as part. (This list is not exhaustive.) A continuant cannot have an occurrent as part: use 'participates in'. An occurrent cannot have a continuant as part: use 'has participant'. An immaterial entity cannot have a material entity as part: use 'location of'. An independent continuant cannot have a specifically dependent continuant as part: use 'bearer of'. A specifically dependent continuant cannot have an independent continuant as part: use 'inheres in'. has_part BFO:0000051 chebi_ontology external protein quality spatial uberon has_part false has_part We use the has_part relation to relate complex qualities to more primitive ones. A complex quality is a collection of qualities. The complex quality cannot exist without the sub-qualities. For example, the quality 'swollen' necessarily comes with the qualities of 'protruding' and 'increased size'. has part has part has_part Q1 has_part Q2 if and only if: every instance of Q1 is a quality_of an entity that has_quality some Q2. PATOC:CJM preceded by X preceded_by Y iff: end(Y) before_or_simultaneous_with start(X) x is preceded by y if and only if the time point at which y ends is before or equivalent to the time point at which x starts. Formally: x preceded by y iff ω(y) <= α(x), where α is a function that maps a process to a start point, and ω is a function that maps a process to an end point. An example is: translation preceded_by transcription; aging preceded_by development (not however death preceded_by aging). Where derives_from links classes of continuants, preceded_by links classes of processes. Clearly, however, these two relations are not independent of each other. Thus if cells of type C1 derive_from cells of type C, then any cell division involving an instance of C1 in a given lineage is preceded_by cellular processes involving an instance of C. The assertion P preceded_by P1 tells us something about Ps in general: that is, it tells us something about what happened earlier, given what we know about what happened later. Thus it does not provide information pointing in the opposite direction, concerning instances of P1 in general; that is, that each is such as to be succeeded by some instance of P. Note that an assertion to the effect that P preceded_by P1 is rather weak; it tells us little about the relations between the underlying instances in virtue of which the preceded_by relation obtains. Typically we will be interested in stronger relations, for example in the relation immediately_preceded_by, or in relations which combine preceded_by with a condition to the effect that the corresponding instances of P and P1 share participants, or that their participants are connected by relations of derivation, or (as a first step along the road to a treatment of causality) that the one process in some way affects (for example, initiates or regulates) the other. is preceded by preceded_by http://www.obofoundry.org/ro/#OBO_REL:preceded_by BFO:0000062 is preceded by takes place after uberon preceded_by preceded_by preceded by preceded_by is preceded by SIO:000249 takes place after Allen:precedes precedes x precedes y if and only if the time point at which x ends is before or equivalent to the time point at which y starts. Formally: x precedes y iff ω(x) <= α(y), where α is a function that maps a process to a start point, and ω is a function that maps a process to an end point. BFO:0000063 uberon precedes precedes precedes precedes occurs in b occurs_in c =def b is a process and c is a material entity or immaterial entity& there exists a spatiotemporal region r and b occupies_spatiotemporal_region r.& forall(t) if b exists_at t then c exists_at t & there exist spatial regions s and s’ where & b spatially_projects_onto s at t& c is occupies_spatial_region s’ at t& s is a proper_continuant_part_of s’ at t occurs_in unfolds in unfolds_in BFO:0000066 external occurs_in occurs_in Paraphrase of definition: a relation between a process and an independent continuant, in which the process takes place entirely within the independent continuant occurs in occurs in site of [copied from inverse property 'occurs in'] b occurs_in c =def b is a process and c is a material entity or immaterial entity& there exists a spatiotemporal region r and b occupies_spatiotemporal_region r.& forall(t) if b exists_at t then c exists_at t & there exist spatial regions s and s’ where & b spatially_projects_onto s at t& c is occupies_spatial_region s’ at t& s is a proper_continuant_part_of s’ at t Paraphrase of definition: a relation between an independent continuant and a process, in which the process takes place entirely within the independent continuant contains process x anterior to y iff x is further along the antero-posterior axis than y, towards the head. An antero-posterior axis is an axis that extends through an organism from head end to opposite end of body or tail. x anterior_to y iff x is further along the antero-posterior axis than y, towards the head. An antero-posterior axis is an axis that bisects an organism from head end to opposite end of body or tail: bearer cjm 2009-07-31T02:15:46Z BSPO:0000096 rostral_to spatial uberon anterior_to anterior_to anterior to anterior_to x anterior to y iff x is further along the antero-posterior axis than y, towards the head. An antero-posterior axis is an axis that extends through an organism from head end to opposite end of body or tail. BSPO:cjm x anterior_to y iff x is further along the antero-posterior axis than y, towards the head. An antero-posterior axis is an axis that bisects an organism from head end to opposite end of body or tail: bearer x distal to y iff x is further along the proximo-distal axis than y, towards the appendage tip. A proximo-distal axis extends from tip of an appendage (distal) to where it joins the body (proximal). x distal_to y iff x is further along the proximo-distal axis than y, towards the appendage tip. A proximo-distal axis extends from tip of an appendage (distal) to where it joins the body (proximal). BSPO:0000097 spatial uberon distal_to distal_to distal to distal_to x distal to y iff x is further along the proximo-distal axis than y, towards the appendage tip. A proximo-distal axis extends from tip of an appendage (distal) to where it joins the body (proximal). BSPO:cjm x distal_to y iff x is further along the proximo-distal axis than y, towards the appendage tip. A proximo-distal axis extends from tip of an appendage (distal) to where it joins the body (proximal). x dorsal to y iff x is further along the dorso-ventral axis than y, towards the back. A dorso-ventral axis is an axis that extends through an organism from back (e.g. spinal column) to front (e.g. belly). x dorsal_to y iff x is further along the dorso-ventral axis than y, towards the back. A dorso-ventral axis is an axis that bisects an organism from back (e.g. spinal column) to front (e.g. belly). BSPO:0000098 spatial uberon dorsal_to dorsal_to dorsal to dorsal_to x dorsal to y iff x is further along the dorso-ventral axis than y, towards the back. A dorso-ventral axis is an axis that extends through an organism from back (e.g. spinal column) to front (e.g. belly). BSPO:cjm x dorsal_to y iff x is further along the dorso-ventral axis than y, towards the back. A dorso-ventral axis is an axis that bisects an organism from back (e.g. spinal column) to front (e.g. belly). x posterior to y iff x is further along the antero-posterior axis than y, towards the body/tail. An antero-posterior axis is an axis that extends through an organism from head end to opposite end of body or tail. x posterior_to y iff x is further along the antero-posterior axis than y, towards the body/tail. An antero-posterior axis is an axis that bisects an organism from head end to opposite end of body or tail. BSPO:0000099 caudal to caudal_to spatial uberon posterior_to posterior_to posterior to posterior_to x posterior to y iff x is further along the antero-posterior axis than y, towards the body/tail. An antero-posterior axis is an axis that extends through an organism from head end to opposite end of body or tail. BSPO:cjm x posterior_to y iff x is further along the antero-posterior axis than y, towards the body/tail. An antero-posterior axis is an axis that bisects an organism from head end to opposite end of body or tail. caudal to x proximal to y iff x is closer to the point of attachment with the body than y. x proximal_to y iff x is closer to the point of attachment with the body than y. BSPO:0000100 spatial uberon proximal_to proximal_to The elbow is proximal to the hand, but distal to the shoulder. proximal to proximal_to x proximal to y iff x is closer to the point of attachment with the body than y. BSPO:cjm x proximal_to y iff x is closer to the point of attachment with the body than y. x ventral to y iff x is further along the dorso-ventral axis than y, towards the front. A dorso-ventral axis is an axis that extends through an organism from back (e.g. spinal column) to front (e.g. belly). x ventral_to y iff x is further along the dorso-ventral axis than y, towards the front. A dorso-ventral axis is an axis that bisects an organism from back (e.g. spinal column) to front (e.g. belly). BSPO:0000102 spatial uberon ventral_to ventral_to ventral to ventral_to x ventral to y iff x is further along the dorso-ventral axis than y, towards the front. A dorso-ventral axis is an axis that extends through an organism from back (e.g. spinal column) to front (e.g. belly). BSPO:cjm x ventral_to y iff x is further along the dorso-ventral axis than y, towards the front. A dorso-ventral axis is an axis that bisects an organism from back (e.g. spinal column) to front (e.g. belly). Crosses at an angle that is 90 degrees, or close to 90 degrees. In any non-linear organism, the main axes are rarely perpendicular when taken through the organism as a whole. BSPO:0000104 spatial approximately_perpendicular_to approximately_perpendicular_to approximately perpendicular to Crosses at an angle that is 90 degrees, or close to 90 degrees. In any non-linear organism, the main axes are rarely perpendicular when taken through the organism as a whole. BSPO:cjm On the opposite side from. For example, the left arm is contralateral to the right arm (and the right leg). BSPO:0000106 spatial contralateral_to contralateral_to contralateral to On the opposite side from. For example, the left arm is contralateral to the right arm (and the right leg). BSPO:cjm Further away from the surface of the organism. Thus, the muscular layer is deep to the skin, but superficial to the intestines. BSPO:0000107 uberon deep_to deep_to deep_to Further away from the surface of the organism. Thus, the muscular layer is deep to the skin, but superficial to the intestines. Near the outer surface of the organism. Thus, skin is superficial to the muscle layer. BSPO:0000108 uberon superficial_to superficial_to superficial_to Near the outer surface of the organism. Thus, skin is superficial to the muscle layer. Closer to the left side of the organism. Example: The dorsal fin is right of the left pectoral fin, but is left of the right eye. On the type level: X left of Y <=> every instance x of X is left of some instance y of Y, and there exists some organism o such that x part of o and y part of o. BSPO:0000110 spatial left_of left_of left of Closer to the left side of the organism. Example: The dorsal fin is right of the left pectoral fin, but is left of the right eye. On the type level: X left of Y <=> every instance x of X is left of some instance y of Y, and there exists some organism o such that x part of o and y part of o. BSPO:cjm Closer to the right side of the organism. Example: The dorsal fin is right of the left pectoral fin, but is left of the right eye. On the type level: X left of Y <=> every instance x of X is right of some instance y of Y, and there exists some organism o such that x part of o and y part of o. BSPO:0000111 spatial right_of right_of right of Closer to the right side of the organism. Example: The dorsal fin is right of the left pectoral fin, but is left of the right eye. On the type level: X left of Y <=> every instance x of X is right of some instance y of Y, and there exists some organism o such that x part of o and y part of o. BSPO:cjm Direcly opposite to. i.e. on the opposite side through the axis. BSPO:0000113 spatial opposite_to opposite_to opposite to Direcly opposite to. i.e. on the opposite side through the axis. BSPO:cjm X posterior_side_of Y <=> if Y is subdivided into two anterior and posterior portions, X is part_of the posterior portion. BSPO:0000122 uberon in_posterior_side_of in_posterior_side_of in_posterior_side_of X posterior_side_of Y <=> if Y is subdivided into two anterior and posterior portions, X is part_of the posterior portion. BSPO:PATO_mtg_2009 X anterior_side_of Y <=> if Y is subdivided into two anterior and posterior portions, X is part_of the anterior portion. BSPO:0000123 uberon in_anterior_side_of in_anterior_side_of in_anterior_side_of X anterior_side_of Y <=> if Y is subdivided into two anterior and posterior portions, X is part_of the anterior portion. BSPO:PATO_mtg_2009 X proximal_side_of Y <=> if Y is subdivided into distal and proximal portions, X is part_of the proximal portion. BSPO:0000124 uberon in_proximal_side_of in_proximal_side_of in_proximal_side_of X proximal_side_of Y <=> if Y is subdivided into distal and proximal portions, X is part_of the proximal portion. BSPO:PATO_mtg_2009 X distal_side_of Y <=> if Y is subdivided into distal and proximal portions, X is part_of the distal portion. BSPO:0000125 uberon in_distal_side_of in_distal_side_of in_distal_side_of X distal_side_of Y <=> if Y is subdivided into distal and proximal portions, X is part_of the distal portion. BSPO:PATO_mtg_2009 X in_lateral_side_of Y <=> if X is in_left_side_of Y or X is in_right_side_of Y. X is often, but not always a paired structure BSPO:0000126 uberon in_lateral_side_of in_lateral_side_of in_lateral_side_of https://github.com/obophenotype/uberon/wiki/Modeling-paired-structures-Design-Pattern X in_lateral_side_of Y <=> if X is in_left_side_of Y or X is in_right_side_of Y. X is often, but not always a paired structure X proximalmost_part_of Y <=> X is part_of Y and X is adjacent_to the proximal boundary of Y BSPO:0001106 uberon proximalmost_part_of proximalmost_part_of proximalmost_part_of X proximalmost_part_of Y <=> X is part_of Y and X is adjacent_to the proximal boundary of Y This relation holds when both the deep_to and ajdacent_to relationship similarly hold. BSPO:0001107 uberon immediately_deep_to immediately_deep_to immediately_deep_to This relation holds when both the deep_to and ajdacent_to relationship similarly hold. BSPO:0015001 spatial finishes_axis finishes_axis A directional axis can be divided by a plane. Following Allen Interval Algebra terminology we say that the side on which the axis initiates starts the axis, with the other side finishing the axis. If s finishes axis x, and x has end point p, then x overlaps p finishes axis crossing at right angles BSPO:0015005 spatial orthogonal_to orthogonal_to orthogonal to crossing at right angles BSPO:cjm Intersects at two points. BSPO:0015006 spatial passes_through passes_through passes through Intersects at two points. BSPO:cjm BSPO:0015007 spatial starts_axis starts_axis A directional axis can be divided by a plane. Following Allen Interval Algebra terminology we say that the side on which the axis initiates starts the axis, with the other side finishing the axis. . If s starts axis x, and x has start point p, then x overlaps p starts axis BSPO:0015008 spatial surface_of surface_of placeholder. used for relation between anatomical surface and an anatomical side. surface of This document is about information artifacts and their representations A (currently) primitive relation that relates an information artifact to an entity. 7/6/2009 Alan Ruttenberg. Following discussion with Jonathan Rees, and introduction of "mentions" relation. Weaken the is_about relationship to be primitive. We will try to build it back up by elaborating the various subproperties that are more precisely defined. Some currently missing phenomena that should be considered "about" are predications - "The only person who knows the answer is sitting beside me" , Allegory, Satire, and other literary forms that can be topical without explicitly mentioning the topic. person:Alan Ruttenberg Smith, Ceusters, Ruttenberg, 2000 years of philosophy is about A person's name denotes the person. A variable name in a computer program denotes some piece of memory. Lexically equivalent strings can denote different things, for instance "Alan" can denote different people. In each case of use, there is a case of the denotation relation obtaining, between "Alan" and the person that is being named. A primitive, instance-level, relation obtaining between an information content entity and some portion of reality. Denotation is what happens when someone creates an information content entity E in order to specifically refer to something. The only relation between E and the thing is that E can be used to 'pick out' the thing. This relation connects those two together. Freedictionary.com sense 3: To signify directly; refer to specifically 2009-11-10 Alan Ruttenberg. Old definition said the following to emphasize the generic nature of this relation. We no longer have 'specifically denotes', which would have been primitive, so make this relation primitive. g denotes r =def r is a portion of reality there is some c that is a concretization of g every c that is a concretization of g specifically denotes r person:Alan Ruttenberg Conversations with Barry Smith, Werner Ceusters, Bjoern Peters, Michel Dumontier, Melanie Courtot, James Malone, Bill Hogan denotes inverse of the relation 'denotes' Person: Jie Zheng, Chris Stoeckert, Mike Conlon denoted by inheres in this fragility inheres in this vase this fragility is a characteristic of this vase this red color inheres in this apple this red color is a characteristic of this apple a relation between a specifically dependent continuant (the characteristic) and any other entity (the bearer), in which the characteristic depends on the bearer for its existence. a relation between a specifically dependent continuant (the dependent) and an independent continuant (the bearer), in which the dependent specifically depends on the bearer for its existence A dependent inheres in its bearer at all times for which the dependent exists. inheres_in Note that this relation was previously called "inheres in", but was changed to be called "characteristic of" because BFO2 uses "inheres in" in a more restricted fashion. This relation differs from BFO2:inheres_in in two respects: (1) it does not impose a range constraint, and thus it allows qualities of processes, as well as of information entities, whereas BFO2 restricts inheres_in to only apply to independent continuants (2) it is declared functional, i.e. something can only be a characteristic of one thing. characteristic of inheres in bearer of this apple is bearer of this red color this vase is bearer of this fragility Inverse of characteristic_of a relation between an independent continuant (the bearer) and a specifically dependent continuant (the dependent), in which the dependent specifically depends on the bearer for its existence A bearer can have many dependents, and its dependents can exist for different periods of time, but none of its dependents can exist when the bearer does not exist. bearer_of is bearer of RO:0000053 uberon bearer_of bearer_of bearer of bearer of has characteristic participates in this blood clot participates in this blood coagulation this input material (or this output material) participates in this process this investigator participates in this investigation a relation between a continuant and a process, in which the continuant is somehow involved in the process participates_in RO:0000056 protein uberon participates_in false participates_in participates in participates in participates_in has participant this blood coagulation has participant this blood clot this investigation has participant this investigator this process has participant this input material (or this output material) a relation between a process and a continuant, in which the continuant is somehow involved in the process Has_participant is a primitive instance-level relation between a process, a continuant, and a time at which the continuant participates in some way in the process. The relation obtains, for example, when this particular process of oxygen exchange across this particular alveolar membrane has_participant this particular sample of hemoglobin at this particular time. has_participant http://www.obofoundry.org/ro/#OBO_REL:has_participant has participant this catalysis function is a function of this enzyme a relation between a function and an independent continuant (the bearer), in which the function specifically depends on the bearer for its existence A function inheres in its bearer at all times for which the function exists, however the function need not be realized at all the times that the function exists. function_of is function of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. function of this red color is a quality of this apple a relation between a quality and an independent continuant (the bearer), in which the quality specifically depends on the bearer for its existence A quality inheres in its bearer at all times for which the quality exists. is quality of quality_of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. quality of this investigator role is a role of this person a relation between a role and an independent continuant (the bearer), in which the role specifically depends on the bearer for its existence A role inheres in its bearer at all times for which the role exists, however the role need not be realized at all the times that the role exists. is role of role_of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. role of this enzyme has function this catalysis function (more colloquially: this enzyme has this catalysis function) a relation between an independent continuant (the bearer) and a function, in which the function specifically depends on the bearer for its existence A bearer can have many functions, and its functions can exist for different periods of time, but none of its functions can exist when the bearer does not exist. A function need not be realized at all the times that the function exists. has_function has function this apple has quality this red color a relation between an independent continuant (the bearer) and a quality, in which the quality specifically depends on the bearer for its existence A bearer can have many qualities, and its qualities can exist for different periods of time, but none of its qualities can exist when the bearer does not exist. has_quality RO:0000086 protein has_quality false has_quality has quality has_quality this person has role this investigator role (more colloquially: this person has this role of investigator) a relation between an independent continuant (the bearer) and a role, in which the role specifically depends on the bearer for its existence A bearer can have many roles, and its roles can exist for different periods of time, but none of its roles can exist when the bearer does not exist. A role need not be realized at all the times that the role exists. has_role RO:0000087 chebi_ontology has_role false false has_role has role has role a relation between an independent continuant (the bearer) and a disposition, in which the disposition specifically depends on the bearer for its existence has disposition inverse of has disposition This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. disposition of is location of my head is the location of my brain this cage is the location of this rat a relation between two independent continuants, the location and the target, in which the target is entirely within the location Most location relations will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ location_of RO:0001015 uberon location_of location_of location of location_of contained in Containment is location not involving parthood, and arises only where some immaterial continuant is involved. Containment obtains in each case between material and immaterial continuants, for instance: lung contained_in thoracic cavity; bladder contained_in pelvic cavity. Hence containment is not a transitive relation. If c part_of c1 at t then we have also, by our definition and by the axioms of mereology applied to spatial regions, c located_in c1 at t. Thus, many examples of instance-level location relations for continuants are in fact cases of instance-level parthood. For material continuants location and parthood coincide. Containment is location not involving parthood, and arises only where some immaterial continuant is involved. To understand this relation, we first define overlap for continuants as follows: c1 overlap c2 at t =def for some c, c part_of c1 at t and c part_of c2 at t. The containment relation on the instance level can then be defined (see definition): Intended meaning: domain: material entity range: spatial region or site (immaterial continuant) contained_in contained in contains RO:0001019 uberon contains contains contains contains located in my brain is located in my head this rat is located in this cage a relation between two independent continuants, the target and the location, in which the target is entirely within the location Location as a relation between instances: The primitive instance-level relation c located_in r at t reflects the fact that each continuant is at any given time associated with exactly one spatial region, namely its exact location. Following we can use this relation to define a further instance-level location relation - not between a continuant and the region which it exactly occupies, but rather between one continuant and another. c is located in c1, in this sense, whenever the spatial region occupied by c is part_of the spatial region occupied by c1. Note that this relation comprehends both the relation of exact location between one continuant and another which obtains when r and r1 are identical (for example, when a portion of fluid exactly fills a cavity), as well as those sorts of inexact location relations which obtain, for example, between brain and head or between ovum and uterus Most location relations will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ located_in http://www.obofoundry.org/ro/#OBO_REL:located_in RO:0001025 protein uberon located_in located_in located in located_in This is redundant with the more specific 'independent and not spatial region' constraint. We leave in the redundant axiom for use with reasoners that do not use negation. This is redundant with the more specific 'independent and not spatial region' constraint. We leave in the redundant axiom for use with reasoners that do not use negation. the surface of my skin is a 2D boundary of my body a relation between a 2D immaterial entity (the boundary) and a material entity, in which the boundary delimits the material entity A 2D boundary may have holes and gaps, but it must be a single connected entity, not an aggregate of several disconnected parts. Although the boundary is two-dimensional, it exists in three-dimensional space and thus has a 3D shape. 2D_boundary_of boundary of is 2D boundary of is boundary of RO:0002000 uberon boundary_of boundary_of 2D boundary of boundary of my body has 2D boundary the surface of my skin a relation between a material entity and a 2D immaterial entity (the boundary), in which the boundary delimits the material entity A 2D boundary may have holes and gaps, but it must be a single connected entity, not an aggregate of several disconnected parts. Although the boundary is two-dimensional, it exists in three-dimensional space and thus has a 3D shape. David Osumi-Sutherland has boundary has_2D_boundary RO:0002002 uberon has_boundary has_boundary has 2D boundary has boundary RO:0002007 uberon bounding_layer_of bounding_layer_of A relationship that applies between a continuant and its outer, bounding layer. Examples include the relationship between a multicellular organism and its integument, between an animal cell and its plasma membrane, and between a membrane bound organelle and its outer/bounding membrane. bounding layer of A relation that holds between two linear structures that are approximately parallel to each other for their entire length and where either the two structures are adjacent to each other or one is part of the other. Note from NCEAS meeting: consider changing primary label David Osumi-Sutherland Example: if we define region of chromosome as any subdivision of a chromosome along its long axis, then we can define a region of chromosome that contains only gene x as 'chromosome region' that coincident_with some 'gene x', where the term gene X corresponds to a genomic sequence. coincident with A 'has regulatory component activity' B if A and B are GO molecular functions (GO_0003674), A has_component B and A is regulated by B. dos 2017-05-24T09:30:46Z has regulatory component activity A relationship that holds between a GO molecular function and a component of that molecular function that negatively regulates the activity of the whole. More formally, A 'has regulatory component activity' B iff :A and B are GO molecular functions (GO_0003674), A has_component B and A is negatively regulated by B. dos 2017-05-24T09:31:01Z By convention GO molecular functions are classified by their effector function. Internal regulatory functions are treated as components. For example, NMDA glutmate receptor activity is a cation channel activity with positive regulatory component 'glutamate binding' and negative regulatory components including 'zinc binding' and 'magnesium binding'. has negative regulatory component activity A relationship that holds between a GO molecular function and a component of that molecular function that positively regulates the activity of the whole. More formally, A 'has regulatory component activity' B iff :A and B are GO molecular functions (GO_0003674), A has_component B and A is positively regulated by B. dos 2017-05-24T09:31:17Z By convention GO molecular functions are classified by their effector function and internal regulatory functions are treated as components. So, for example calmodulin has a protein binding activity that has positive regulatory component activity calcium binding activity. Receptor tyrosine kinase activity is a tyrosine kinase activity that has positive regulatory component 'ligand binding'. has positive regulatory component activity dos 2017-05-24T09:44:33Z A 'has component activity' B if A is A and B are molecular functions (GO_0003674) and A has_component B. has component activity w 'has process component' p if p and w are processes, w 'has part' p and w is such that it can be directly disassembled into into n parts p, p2, p3, ..., pn, where these parts are of similar type. dos 2017-05-24T09:49:21Z has component process dos 2017-09-17T13:52:24Z Process(P2) is directly regulated by process(P1) iff: P1 regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding regulates the kinase activity (P2) of protein B then P1 directly regulates P2. directly regulated by Process(P2) is directly regulated by process(P1) iff: P1 regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding regulates the kinase activity (P2) of protein B then P1 directly regulates P2. GOC:dos Process(P2) is directly negatively regulated by process(P1) iff: P1 negatively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding negatively regulates the kinase activity (P2) of protein B then P2 directly negatively regulated by P1. dos 2017-09-17T13:52:38Z directly negatively regulated by Process(P2) is directly negatively regulated by process(P1) iff: P1 negatively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding negatively regulates the kinase activity (P2) of protein B then P2 directly negatively regulated by P1. GOC:dos Process(P2) is directly postively regulated by process(P1) iff: P1 positively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding positively regulates the kinase activity (P2) of protein B then P2 is directly postively regulated by P1. dos 2017-09-17T13:52:47Z directly positively regulated by Process(P2) is directly postively regulated by process(P1) iff: P1 positively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding positively regulates the kinase activity (P2) of protein B then P2 is directly postively regulated by P1. GOC:dos A 'has effector activity' B if A and B are GO molecular functions (GO_0003674), A 'has component activity' B and B is the effector (output function) of B. Each compound function has only one effector activity. dos 2017-09-22T14:14:36Z This relation is designed for constructing compound molecular functions, typically in combination with one or more regulatory component activity relations. has effector activity A 'has effector activity' B if A and B are GO molecular functions (GO_0003674), A 'has component activity' B and B is the effector (output function) of B. Each compound function has only one effector activity. GOC:dos David Osumi-Sutherland X ends_after Y iff: end(Y) before_or_simultaneous_with end(X) ends after X immediately_preceded_by Y iff: end(X) simultaneous_with start(Y) David Osumi-Sutherland starts_at_end_of A non-transitive temporal relation in which one process immediately precedes another process, such that there is no interval of time between the two processes[SIO:000251]. RO:0002087 directly preceded by is directly preceded by is immediately preceded by starts_at_end_of uberon immediately_preceded_by immediately_preceded_by X immediately_preceded_by Y iff: end(X) simultaneous_with start(Y) immediately preceded by immediately_preceded_by A non-transitive temporal relation in which one process immediately precedes another process, such that there is no interval of time between the two processes[SIO:000251]. SIO:000251 is immediately preceded by SIO:000251 David Osumi-Sutherland ends_at_start_of meets X immediately_precedes_Y iff: end(X) simultaneous_with start(Y) immediately precedes x overlaps y if and only if there exists some z such that x has part z and z part of y x overlaps y iff they have some part in common. http://purl.obolibrary.org/obo/BFO_0000051 some (http://purl.obolibrary.org/obo/BFO_0000050 some ?Y) RO:0002131 spatial overlaps overlaps "(forall (x y) (iff (overlaps x y) (exists (z) (and (part of z x) (part of z y)))))" CLIF [] overlaps overlaps true x overlaps y iff they have some part in common. BSPO:cjm X continuous_with Y if and only if X and Y share a fiat boundary. David Osumi-Sutherland connected to The label for this relation was previously connected to. I relabeled this to "continuous with". The standard notion of connectedness does not imply shared boundaries - e.g. Glasgow connected_to Edinburgh via M8; my patella connected_to my femur (via patellar-femoral joint) RO:0002150 uberon continuous_with continuous_with continuous with continuous_with FMA:85972 lactation SubClassOf 'only in taxon' some 'Mammalia' S only_in_taxon T iff: S SubClassOf in_taxon only T U only_in_taxon T: U is a feature found in only in organisms of species of taxon T. The feature cannot be found in an organism of any species outside of (not subsumed by) that taxon. Down-propagates in U hierarchy, up-propagates in T hierarchy (species taxonomy). Implies applicable_to_taxon. x only in taxon y if and only if x is in taxon y, and there is no other organism z such that y!=z a and x is in taxon z. The original intent was to treat this as a macro that expands to 'in taxon' only ?Y - however, this is not necessary if we instead have supplemental axioms that state that each pair of sibling tax have a disjointness axiom using the 'in taxon' property - e.g. 'in taxon' some Eukaryota DisjointWith 'in taxon' some Eubacteria Chris Mungall RO:0002160 never_outside_taxon specific_to specific_to_taxon protein uberon only_in_taxon false only_in_taxon Down-propagates. The original name for this in the paper is 'specific_to'. Applicable to genes because some genes are lost in sub-species (strains) of a species. only in taxon only_in_taxon S only_in_taxon T iff: S SubClassOf in_taxon only T U only_in_taxon T: U is a feature found in only in organisms of species of taxon T. The feature cannot be found in an organism of any species outside of (not subsumed by) that taxon. Down-propagates in U hierarchy, up-propagates in T hierarchy (species taxonomy). Implies applicable_to_taxon. ROC:Waclaw x is in taxon y if an only if y is an organism, and the relationship between x and y is one of: part of (reflexive), developmentally preceded by, derives from, secreted by, expressed. Chris Mungall Jennifer Deegan RO:0002162 uberon in_taxon in_taxon Connects a biological entity to its taxon of origin. in taxon A is spatially_disjoint_from B if and only if they have no parts in common There are two ways to encode this as a shortcut relation. The other possibility to use an annotation assertion between two classes, and expand this to a disjointness axiom. Chris Mungall Note that it would be possible to use the relation to label the relationship between a near infinite number of structures - between the rings of saturn and my left earlobe. The intent is that this is used for parsiomoniously for disambiguation purposes - for example, between siblings in a jointly exhaustive pairwise disjointness hierarchy BFO_0000051 exactly 0 (BFO_0000050 some ?Y) spatially disjoint from https://github.com/obophenotype/uberon/wiki/Part-disjointness-Design-Pattern a 'toe distal phalanx bone' that is connected to a 'toe medial phalanx bone' (an interphalangeal joint *connects* these two bones). Binary relationship: x connected_to y if and only if there exists some z such that z connects x and y in a ternary connected_to(x,y,z) relationship. a is connected to b if and only if a and b are discrete structure, and there exists some connecting structure c, such that c connects a and b RO:0002170 uberon connected_to connected_to Connection does not imply overlaps. connected to connected to https://github.com/obophenotype/uberon/wiki/Connectivity-Design-Pattern https://github.com/obophenotype/uberon/wiki/Modeling-articulations-Design-Pattern Binary relationship: x connected_to y if and only if there exists some z such that z connects x and y in a ternary connected_to(x,y,z) relationship. Binary relationship: z connects x if and only if there exists some y such that z connects x and y in a ternary connected_to(x,y,z) relationship. this is currently used for both structural relationships (such as between a valve and the chamber it connects) and abstract relationships (anatomical lines and the entities they connect) RO:0002176 uberon connects connects connects Binary relationship: z connects x if and only if there exists some y such that z connects x and y in a ternary connected_to(x,y,z) relationship. a is attached to part of b if a is attached to b, or a is attached to some p, where p is part of b. attached to part of (anatomical structure to anatomical structure) RO:0002177 uberon attaches_to_part_of attaches_to_part_of attached to part of attaches_to_part_of true true RO:0002179 drains blood from drains from uberon drains drains source: Wikipedia drains w 'has component' p if w 'has part' p and w is such that it can be directly disassembled into into n parts p, p2, p3, ..., pn, where these parts are of similar type. The definition of 'has component' is still under discussion. The challenge is in providing a definition that does not imply transitivity. For use in recording has_part with a cardinality constraint, because OWL does not permit cardinality constraints to be used in combination with transitive object properties. In situations where you would want to say something like 'has part exactly 5 digit, you would instead use has_component exactly 5 digit. RO:0002180 protein uberon has_component false has_component has component has component has_component x develops from y if and only if either (a) x directly develops from y or (b) there exists some z such that x directly develops from z and z develops from y Chris Mungall David Osumi-Sutherland Melissa Haendel Terry Meehan RO:0002202 uberon develops_from develops_from This is the transitive form of the develops from relation develops from develops_from inverse of develops from Chris Mungall David Osumi-Sutherland Terry Meehan RO:0002203 uberon develops_into develops_into develops into develops_into process(P1) regulates process(P2) iff: P1 results in the initiation or termination of P2 OR affects the frequency of its initiation or termination OR affects the magnitude or rate of output of P2. We use 'regulates' here to specifically imply control. However, many colloquial usages of the term correctly correspond to the weaker relation of 'causally upstream of or within' (aka influences). Consider relabeling to make things more explicit Chris Mungall David Hill Tanya Berardini GO Regulation precludes parthood; the regulatory process may not be within the regulated process. regulates (processual) false RO:0002211 external regulates regulates regulates regulates Process(P1) negatively regulates process(P2) iff: P1 terminates P2, or P1 descreases the the frequency of initiation of P2 or the magnitude or rate of output of P2. Chris Mungall negatively regulates (process to process) RO:0002212 external negatively_regulates negatively_regulates negatively regulates negatively regulates Process(P1) postively regulates process(P2) iff: P1 initiates P2, or P1 increases the the frequency of initiation of P2 or the magnitude or rate of output of P2. Chris Mungall positively regulates (process to process) RO:0002213 external positively_regulates positively_regulates positively regulates positively regulates mechanosensory neuron capable of detection of mechanical stimulus involved in sensory perception (GO:0050974) osteoclast SubClassOf 'capable of' some 'bone resorption' A relation between a material entity (such as a cell) and a process, in which the material entity has the ability to carry out the process. Chris Mungall has function realized in For compatibility with BFO, this relation has a shortcut definition in which the expression "capable of some P" expands to "bearer_of (some realized_by only P)". RO:0002215 protein uberon capable_of false capable_of capable of capable of capable_of c stands in this relationship to p if and only if there exists some p' such that c is capable_of p', and p' is part_of p. Chris Mungall has function in RO:0002216 uberon capable_of_part_of capable_of_part_of capable of part of capable of part of true x surrounded_by y if and only if (1) x is adjacent to y and for every region r that is adjacent to x, r overlaps y (2) the shared boundary between x and y occupies the majority of the outermost boundary of x x surrounded_by y iff: x is adjacent to y and for every region r adjacent to x, r overlaps y Chris Mungall RO:0002219 uberon surrounded_by surrounded_by surrounded by surrounded_by x surrounded_by y iff: x is adjacent to y and for every region r adjacent to x, r overlaps y A caterpillar walking on the surface of a leaf is adjacent_to the leaf, if one of the caterpillar appendages is touching the leaf. In contrast, a butterfly flying close to a flower is not considered adjacent, unless there are any touching parts. The epidermis layer of a vertebrate is adjacent to the dermis. The plasma membrane of a cell is adjacent to the cytoplasm, and also to the cell lumen which the cytoplasm occupies. The skin of the forelimb is adjacent to the skin of the torso if these are considered anatomical subdivisions with a defined border. Otherwise a relation such as continuous_with would be used. x adjacent to y if and only if x and y share a boundary. x adjacent_to y iff: x and y share a boundary This relation acts as a join point with BSPO Chris Mungall RO:0002220 spatial uberon adjacent_to adjacent_to adjacent to adjacent to adjacent_to A caterpillar walking on the surface of a leaf is adjacent_to the leaf, if one of the caterpillar appendages is touching the leaf. In contrast, a butterfly flying close to a flower is not considered adjacent, unless there are any touching parts. inverse of surrounded by inverse of surrounded_by Chris Mungall RO:0002221 uberon surrounds surrounds surrounds surrounds inverse of surrounded_by Chris Mungall Do not use this relation directly. It is ended as a grouping for relations between occurrents involving the relative timing of their starts and ends. https://docs.google.com/document/d/1kBv1ep_9g3sTR-SD3jqzFqhuwo9TPNF-l-9fUDbO6rM/edit?pli=1 A relation that holds between two occurrents. This is a grouping relation that collects together all the Allen relations. temporally related to Relation between occurrents, shares a start boundary with. RO:0002223 uberon starts starts starts Relation between occurrents, shares a start boundary with. Allen:starts RO:0002224 uberon starts_with starts_with starts with Relation between occurrents, shares an end boundary with. RO:0002229 finishes uberon ends ends ends Relation between occurrents, shares an end boundary with. Allen:starts ZFS:finishes RO:0002230 uberon ends_with ends_with ends with p has input c iff: p is a process, c is a material entity, c is a participant in p, c is present at the start of p, and the state of c is modified during p. Chris Mungall consumes has input Mammalian thymus has developmental contribution from some pharyngeal pouch 3; Mammalian thymus has developmental contribution from some pharyngeal pouch 4 [Kardong] x has developmental contribution from y iff x has some part z such that z develops from y Chris Mungall RO:0002254 uberon has_developmental_contribution_from has_developmental_contribution_from has developmental contribution from has developmental contribution from inverse of has developmental contribution from Chris Mungall RO:0002255 uberon developmentally_contributes_to developmentally_contributes_to developmentally contributes to developmentally_contributes_to t1 developmentally_induced_by t2 if there is a process of organ induction (GO:0001759) with t1 and t2 as interacting participants. t2 causes t1 to change its fate from a precursor tissue type T to T', where T' develops_from T. RO:0002256 uberon developmentally_induced_by developmentally_induced_by sources for developmentally_induced_by relationships in Uberon: Developmental Biology, Gilbert, 8th edition, figure 6.5(F) developmentally_induced_by t1 developmentally_induced_by t2 if there is a process of organ induction (GO:0001759) with t1 and t2 as interacting participants. t2 causes t1 to change its fate from a precursor tissue type T to T', where T' develops_from T. GO:0001759 Candidate definition: x developmentally related to y if and only if there exists some developmental process (GO:0032502) p such that x and y both participates in p, and x is the output of p and y is the input of p false Chris Mungall In general you should not use this relation to make assertions - use one of the more specific relations below this one This relation groups together various other developmental relations. It is fairly generic, encompassing induction, developmental contribution and direct and transitive develops from developmentally preceded by A faulty traffic light (material entity) whose malfunctioning (a process) is causally upstream of a traffic collision (a process): the traffic light acts upstream of the collision. c acts upstream of p if and only if c enables some f that is involved in p' and p' occurs chronologically before p, is not part of p, and affects the execution of p. c is a material entity and f, p, p' are processes. acts upstream of A gene product that has some activity, where that activity may be a part of a pathway or upstream of the pathway. c acts upstream of or within p if c is enables f, and f is causally upstream of or within p. c is a material entity and p is an process. affects acts upstream of or within RO:0002285 uberon developmentally_replaces developmentally_replaces developmentally_replaces Inverse of developmentally preceded by Chris Mungall developmentally succeeded by cjm holds between x and y if and only if x is causally upstream of y and the progression of x increases the frequency, rate or extent of y causally upstream of, positive effect cjm holds between x and y if and only if x is causally upstream of y and the progression of x decreases the frequency, rate or extent of y causally upstream of, negative effect q characteristic of part of w if and only if there exists some p such that q inheres in p and p part of w. q inheres in part of w if and only if there exists some p such that q inheres in p and p part of w. Because part_of is transitive, inheres in is a sub-relation of characteristic of part of Because part_of is transitive, inheres in is a sub-relation of inheres in part of Chris Mungall inheres in part of characteristic of part of inheres in part of true A relationship that holds via some environmental process Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving the process of evolution. evolutionarily related to A mereological relationship or a topological relationship Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving parthood or connectivity relationships mereotopologically related to A relationship that holds between entities participating in some developmental process (GO:0032502) Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving organismal development developmentally related to a particular instances of akt-2 enables some instance of protein kinase activity Chris Mungall catalyzes executes has is catalyzing is executing This relation differs from the parent relation 'capable of' in that the parent is weaker and only expresses a capability that may not be actually realized, whereas this relation is always realized. This relation is currently used experimentally by the Gene Ontology Consortium. It may not be stable and may be obsoleted at some future time. enables A grouping relationship for any relationship directly involving a function, or that holds because of a function of one of the related entities. Chris Mungall This is a grouping relation that collects relations used for the purpose of connecting structure and function RO:0002328 uberon functionally_related_to functionally_related_to functionally related to functionally related to this relation holds between c and p when c is part of some c', and c' is capable of p. Chris Mungall false RO:0002329 uberon part_of_structure_that_is_capable_of part_of_structure_that_is_capable_of part of structure that is capable of part of structure that is capable of true c involved_in p if and only if c enables some process p', and p' is part of p Chris Mungall actively involved in enables part of involved in inverse of enables Chris Mungall enabled by inverse of regulates Chris Mungall regulated by (processual) regulated by inverse of negatively regulates Chris Mungall negatively regulated by inverse of positively regulates Chris Mungall positively regulated by An organism that is a member of a population of organisms is member of is a mereological relation between a item and a collection. is member of member part of SIO RO:0002350 uberon member_of member_of member of member of has member is a mereological relation between a collection and an item. SIO RO:0002351 uberon has_member has_member has member has member inverse of has input Chris Mungall input of RO:0002353 protein output_of false output_of output_of a is attached to b if and only if a and b are discrete objects or object parts, and there are physical connections between a and b such that a force pulling a will move b, or a force pulling b will move a Chris Mungall attached to (anatomical structure to anatomical structure) RO:0002371 uberon attaches_to attaches_to attached to attaches_to m has_muscle_origin s iff m is attached_to s, and it is the case that when m contracts, s does not move. The site of the origin tends to be more proximal and have greater mass than what the other end attaches to. m has_muscle_origin s iff m is attaches_to s, and it is the case that when m contracts, s does not move. The site of the origin tends to be more proximal and have greater mass than what the other end attaches to. Chris Mungall Wikipedia:Insertion_(anatomy) RO:0002372 uberon has_muscle_origin has_muscle_origin The origin is the end of a muscle that attaches to the more fixed part of the skeleton, which is the proximal end in limb muscles has muscle origin has_muscle_origin We need to import uberon muscle to create a stricter domain constraint m has_muscle_origin s iff m is attaches_to s, and it is the case that when m contracts, s does not move. The site of the origin tends to be more proximal and have greater mass than what the other end attaches to. m has_muscle_insertion s iff m is attaches_to s, and it is the case that when m contracts, s moves. Insertions are usually connections of muscle via tendon to bone. RO:0002373 uberon has_muscle_insertion has_muscle_insertion The insertion is the point of attachment of a muscle that moves the most when the muscle shortens, or the most distal end of limb muscles has_muscle_insertion m has_muscle_insertion s iff m is attaches_to s, and it is the case that when m contracts, s moves. Insertions are usually connections of muscle via tendon to bone. RO:0002376 uberon drains into tributary_of tributary_of tributary_of drains into dbpowl:drainsTo A lump of clay and a statue x spatially_coextensive_with y if and inly if x and y have the same location Chris Mungall This relation is added for formal completeness. It is unlikely to be used in many practical scenarios spatially coextensive with we need to check if FMA branch_of implies part_of. the relation we intend to use here should - for example, see vestibulocochlear nerve RO:0002380 uberon branching_part_of branching_part_of branching_part_of x has developmental potential involving y iff x is capable of a developmental process with output y. y may be the successor of x, or may be a different structure in the vicinity (as for example in the case of developmental induction). Chris Mungall has developmental potential involving x has potential to developmentrally contribute to y iff x developmentally contributes to y or x is capable of developmentally contributing to y x has potential to developmentrally contribute to y iff x developmentally contributes to y or x is capable of developmentally contributing to y Chris Mungall RO:0002385 uberon has_potential_to_developmentally_contribute_to has_potential_to_developmentally_contribute_to has potential to developmentally contribute to has potential to developmentally contribute to x has the potential to develop into y iff x develops into y or if x is capable of developing into y x has the potential to develop into y iff x develops into y or if x is capable of developing into y Chris Mungall RO:0002387 uberon has_potential_to_develop_into has_potential_to_develop_into has potential to develop into has potential to develop into x has potential to directly develop into y iff x directly develops into y or x is capable of directly developing into y Chris Mungall has potential to directly develop into inverse of upstream of Chris Mungall causally downstream of Chris Mungall immediately causally downstream of This relation groups causal relations between material entities and causal relations between processes This branch of the ontology deals with causal relations between entities. It is divided into two branches: causal relations between occurrents/processes, and causal relations between material entities. We take an 'activity flow-centric approach', with the former as primary, and define causal relations between material entities in terms of causal relations between occurrents. To define causal relations in an activity-flow type network, we make use of 3 primitives: * Temporal: how do the intervals of the two occurrents relate? * Is the causal relation regulatory? * Is the influence positive or negative The first of these can be formalized in terms of the Allen Interval Algebra. Informally, the 3 bins we care about are 'direct', 'indirect' or overlapping. Note that all causal relations should be classified under a RO temporal relation (see the branch under 'temporally related to'). Note that all causal relations are temporal, but not all temporal relations are causal. Two occurrents can be related in time without being causally connected. We take causal influence to be primitive, elucidated as being such that has the upstream changed, some qualities of the donwstream would necessarily be modified. For the second, we consider a relationship to be regulatory if the system in which the activities occur is capable of altering the relationship to achieve some objective. This could include changing the rate of production of a molecule. For the third, we consider the effect of the upstream process on the output(s) of the downstream process. If the level of output is increased, or the rate of production of the output is increased, then the direction is increased. Direction can be positive, negative or neutral or capable of either direction. Two positives in succession yield a positive, two negatives in succession yield a positive, otherwise the default assumption is that the net effect is canceled and the influence is neutral. Each of these 3 primitives can be composed to yield a cross-product of different relation types. Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causally related to p is causally upstream of q if and only if p precedes q and p and q are linked in a causal chain Chris Mungall causally upstream of p is immediately causally upstream of q iff both (a) p immediately precedes q and (b) p is causally upstream of q. In addition, the output of p must be an input of q. Chris Mungall immediately causally upstream of p 'causally upstream or within' q iff (1) the end of p is before the end of q and (2) the execution of p exerts some causal influence over the outputs of q; i.e. if p was abolished or the outputs of p were to be modified, this would necessarily affect q. We would like to make this disjoint with 'preceded by', but this is prohibited in OWL2 Chris Mungall influences (processual) affects causally upstream of or within inverse of causally upstream of or within Chris Mungall causally downstream of or within c involved in regulation of p if c is involved in some p' and p' regulates some p Chris Mungall involved in regulation of c involved in regulation of p if c is involved in some p' and p' positively regulates some p Chris Mungall involved in positive regulation of c involved in regulation of p if c is involved in some p' and p' negatively regulates some p Chris Mungall involved in negative regulation of c involved in or regulates p if and only if either (i) c is involved in p or (ii) c is involved in regulation of p OWL does not allow defining object properties via a Union Chris Mungall involved in or reguates involved in or involved in regulation of A protein that enables activity in a cytosol. c executes activity in d if and only if c enables p and p occurs_in d. Assuming no action at a distance by gene products, if a gene product enables (is capable of) a process that occurs in some structure, it must have at least some part in that structure. Chris Mungall executes activity in enables activity in is active in true c executes activity in d if and only if c enables p and p occurs_in d. Assuming no action at a distance by gene products, if a gene product enables (is capable of) a process that occurs in some structure, it must have at least some part in that structure. GOC:cjm GOC:dos RO:0002433 uberon contributes_to_morphology_of contributes_to_morphology_of contributes to morphology of A relationship that holds between two entities in which the processes executed by the two entities are causally connected. Considering relabeling as 'pairwise interacts with' This relation and all sub-relations can be applied to either (1) pairs of entities that are interacting at any moment of time (2) populations or species of entity whose members have the disposition to interact (3) classes whose members have the disposition to interact. Chris Mungall Note that this relationship type, and sub-relationship types may be redundant with process terms from other ontologies. For example, the symbiotic relationship hierarchy parallels GO. The relations are provided as a convenient shortcut. Consider using the more expressive processual form to capture your data. In the future, these relations will be linked to their cognate processes through rules. in pairwise interaction with interacts with http://purl.obolibrary.org/obo/MI_0914 http://purl.obolibrary.org/obo/ro/docs/interaction-relations/ http://purl.obolibrary.org/obo/MI_0914 https://github.com/oborel/obo-relations/wiki/InteractionRelations An interaction relationship in which the two partners are molecular entities that directly physically interact with each other for example via a stable binding interaction or a brief interaction during which one modifies the other. Chris Mungall binds molecularly binds with molecularly interacts with http://purl.obolibrary.org/obo/MI_0915 http://purl.obolibrary.org/obo/MI_0915 Axiomatization to GO to be added later Chris Mungall An interaction relation between x and y in which x catalyzes a reaction in which a phosphate group is added to y. phosphorylates The entity A, immediately upstream of the entity B, has an activity that regulates an activity performed by B. For example, A and B may be gene products and binding of B by A regulates the kinase activity of B. A and B can be physically interacting but not necessarily. Immediately upstream means there are no intermediate entity between A and B. Chris Mungall Vasundra Touré molecularly controls directly regulates activity of The entity A, immediately upstream of the entity B, has an activity that negatively regulates an activity performed by B. For example, A and B may be gene products and binding of B by A negatively regulates the kinase activity of B. Chris Mungall Vasundra Touré directly inhibits molecularly decreases activity of directly negatively regulates activity of The entity A, immediately upstream of the entity B, has an activity that positively regulates an activity performed by B. For example, A and B may be gene products and binding of B by A positively regulates the kinase activity of B. Chris Mungall Vasundra Touré directly activates molecularly increases activity of directly positively regulates activity of Chris Mungall This property or its subproperties is not to be used directly. These properties exist as helper properties that are used to support OWL reasoning. helper property (not for use in curation) x composed_primarily_of y iff: more than half of the mass of x is made from parts of y RO:0002473 uberon RO:0002473 composed_primarily_of composed_primarily_of composed primarily of p has part that occurs in c if and only if there exists some p1, such that p has_part p1, and p1 occurs in c. Chris Mungall has part that occurs in true Chris Mungall is kinase activity Relation between continuant c and occurrent s, such that every instance of c comes into existing during some s. RO:0002488 begins_to_exist_during uberon RO:0002488 existence_starts_during existence_starts_during existence starts during Relation between continuant c and occurrent s, such that every instance of c comes into existing during some s. Relation between continuant and occurrent, such that c comes into existence at the start of p. RO:0002489 uberon existence_starts_with existence_starts_with existence starts with Relation between continuant and occurrent, such that c comes into existence at the start of p. RO:0002491 uberon existence_starts_and_ends_during existence_starts_and_ends_during existence starts and ends during Relation between continuant c and occurrent s, such that every instance of c ceases to exist during some s, if it does not die prematurely. RO:0002492 ceases_to_exist_during uberon RO:0002492 existence_ends_during existence_ends_during existence ends during Relation between continuant c and occurrent s, such that every instance of c ceases to exist during some s, if it does not die prematurely. Relation between continuant and occurrent, such that c ceases to exist at the end of p. RO:0002493 uberon existence_ends_with existence_ends_with existence ends with Relation between continuant and occurrent, such that c ceases to exist at the end of p. RO:0002494 transforms from uberon transformation_of transformation_of transformation of transforms from SIO:000657 RO:0002495 direct_transformation_of immediately transforms from uberon immediate_transformation_of immediate_transformation_of immediate transformation of direct_transformation_of immediately transforms from SIO:000658 RO:0002496 uberon existence_starts_during_or_after existence_starts_during_or_after existence starts during or after RO:0002497 uberon existence_ends_during_or_before existence_ends_during_or_before existence ends during or before A relationship between a material entity and a process where the material entity has some causal role that influences the process causal agent in process p is causally related to q if and only if p or any part of p and q or any part of q are linked by a chain of events where each event pair is one of direct activation or direct inhibition. p may be upstream, downstream, part of or a container of q. Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causal relation between processes Chris Mungall depends on The intent is that the process branch of the causal property hierarchy is primary (causal relations hold between occurrents/processes), and that the material branch is defined in terms of the process branch Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causal relation between entities A relation between a subdivision of an organism and the single subdivision of skeleton that provides structural support for that subdivision. RO:0002551 uberon has sekeletal support has supporting framework has_skeleton has_skeleton has skeleton A relation between a subdivision of an organism and the single subdivision of skeleton that provides structural support for that subdivision. Chris Mungall causally influenced by (entity-centric) causally influenced by Chris Mungall interaction relation helper property http://purl.obolibrary.org/obo/ro/docs/interaction-relations/ https://github.com/oborel/obo-relations/wiki/InteractionRelations Chris Mungall molecular interaction relation helper property The entity or characteristic A is causally upstream of the entity or characteristic B, A having an effect on B. An entity corresponds to any biological type of entity as long as a mass is measurable. A characteristic corresponds to a particular specificity of an entity (e.g., phenotype, shape, size). Chris Mungall Vasundra Touré causally influences (entity-centric) causally influences A relation that holds between elements of a musculoskeletal system or its analogs. Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving the biomechanical processes. biomechanically related to RO:0002569 uberon has_branching_part has_branching_part has branching part RO:0002572 uberon luminal_space_of luminal_space_of luminal space of RO:0002576 uberon skeleton_of skeleton_of skeleton of Process(P1) directly regulates process(P2) iff: P1 regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding regulates the kinase activity (P2) of protein B then P1 directly regulates P2. Chris Mungall directly regulates (processual) directly regulates gland SubClassOf 'has part structure that is capable of' some 'secretion by cell' s 'has part structure that is capable of' p if and only if there exists some part x such that s 'has part' x and x 'capable of' p Chris Mungall has part structure that is capable of A relationship that holds between a material entity and a process in which causality is involved, with either the material entity or some part of the material entity exerting some influence over the process, or the process influencing some aspect of the material entity. Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. Chris Mungall causal relation between material entity and a process pyrethroid -> growth Holds between c and p if and only if c is capable of some activity a, and a regulates p. capable of regulating Holds between c and p if and only if c is capable of some activity a, and a negatively regulates p. capable of negatively regulating renin -> arteriolar smooth muscle contraction Holds between c and p if and only if c is capable of some activity a, and a positively regulates p. capable of positively regulating Inverse of 'causal agent in process' process has causal agent Process(P1) directly postively regulates process(P2) iff: P1 positively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding positively regulates the kinase activity (P2) of protein B then P1 directly positively regulates P2. directly positively regulates (process to process) directly positively regulates Process(P1) directly negatively regulates process(P2) iff: P1 negatively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding negatively regulates the kinase activity (P2) of protein B then P1 directly negatively regulates P2. directly negatively regulates (process to process) directly negatively regulates a produces b if some process that occurs_in a has_output b, where a and b are material entities. Examples: hybridoma cell line produces monoclonal antibody reagent; chondroblast produces avascular GAG-rich matrix. Melissa Haendel RO:0003000 uberon produces produces Note that this definition doesn't quite distinguish the output of a transformation process from a production process, which is related to the identity/granularity issue. produces produces a produced_by b iff some process that occurs_in b has_output a. Melissa Haendel RO:0003001 uberon produced_by produced_by produced by produced_by Holds between an entity and an process P where the entity enables some larger compound process, and that larger process has-part P. cjm 2018-01-25T23:20:13Z enables subfunction cjm 2018-01-26T23:49:30Z acts upstream of or within, positive effect cjm 2018-01-26T23:49:51Z acts upstream of or within, negative effect c 'acts upstream of, positive effect' p if c is enables f, and f is causally upstream of p, and the direction of f is positive cjm 2018-01-26T23:53:14Z acts upstream of, positive effect c 'acts upstream of, negative effect' p if c is enables f, and f is causally upstream of p, and the direction of f is negative cjm 2018-01-26T23:53:22Z acts upstream of, negative effect cjm 2018-03-13T23:55:05Z causally upstream of or within, negative effect cjm 2018-03-13T23:55:19Z causally upstream of or within, positive effect The entity A has an activity that regulates an activity of the entity B. For example, A and B are gene products where the catalytic activity of A regulates the kinase activity of B. Vasundra Touré regulates activity of x is parallel t y iff x and y are lines or planes which when extended indefinitely do not cross. spatial parallel_to parallel to x is parallel t y iff x and y are lines or planes which when extended indefinitely do not cross. BSPO:curators q1 decreased_in_magnitude_relative_to q2 if and only if magnitude(q1) < magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. quality decreased_in_magnitude_relative_to This relation is used to determine the 'directionality' of relative qualities such as 'decreased strength', relative to the parent type, 'strength'. decreased_in_magnitude_relative_to q1 decreased_in_magnitude_relative_to q2 if and only if magnitude(q1) < magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. PATOC:CJM q1 different_in_magnitude_relative_to q2 if and only if magnitude(q1) NOT =~ magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. quality different_in_magnitude_relative_to different_in_magnitude_relative_to q1 different_in_magnitude_relative_to q2 if and only if magnitude(q1) NOT =~ magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. PATOC:CJM s3 has_cross_section s3 if and only if : there exists some 2d plane that intersects the bearer of s3, and the impression of s3 upon that plane has shape quality s2. quality has_cross_section Example: a spherical object has the quality of being spherical, and the spherical quality has_cross_section round. has_cross_section s3 has_cross_section s3 if and only if : there exists some 2d plane that intersects the bearer of s3, and the impression of s3 upon that plane has shape quality s2. PATOC:CJM q1 increased_in_magnitude_relative_to q2 if and only if magnitude(q1) > magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. quality increased_in_magnitude_relative_to This relation is used to determine the 'directionality' of relative qualities such as 'increased strength', relative to the parent type, 'strength'. increased_in_magnitude_relative_to q1 increased_in_magnitude_relative_to q2 if and only if magnitude(q1) > magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. PATOC:CJM q1 reciprocal_of q2 if and only if : q1 and q2 are relational qualities and a phenotype e q1 e2 mutually implies a phenotype e2 q2 e. quality reciprocal_of There are frequently two ways to state the same thing: we can say 'spermatocyte lacks asters' or 'asters absent from spermatocyte'. In this case the quality is 'lacking all parts of type' - it is a (relational) quality of the spermatocyte, and it is with respect to instances of 'aster'. One of the popular requirements of PATO is that it continue to support 'absent', so we need to relate statements which use this quality to the 'lacking all parts of type' quality. reciprocal_of q1 reciprocal_of q2 if and only if : q1 and q2 are relational qualities and a phenotype e q1 e2 mutually implies a phenotype e2 q2 e. PATOC:CJM q1 similar_in_magnitude_relative_to q2 if and only if magnitude(q1) =~ magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. quality similar_in_magnitude_relative_to similar_in_magnitude_relative_to q1 similar_in_magnitude_relative_to q2 if and only if magnitude(q1) =~ magnitude(q2). Here, magnitude(q) is a function that maps a quality to a unit-invariant scale. PATOC:CJM protein has_gene_template false has_gene_template x anteriorly_connected_to y iff the anterior part of x is connected to y. i.e. x connected_to y and x posterior_to y. uberon anteriorly_connected_to anteriorly connected to x anteriorly_connected_to y iff the anterior part of x is connected to y. i.e. x connected_to y and x posterior_to y. carries uberon channel_for channel for uberon channels_from channels_from x is a conduit for y iff y passes through the lumen of x. uberon conduit_for conduit for Relationship between a fluid and a material entity, where the fluid is the output of a realization of a filtration role that inheres in the material entity. uberon filtered_through Relationship between a fluid and a filtration barrier, where the portion of fluid arises as a transformation of another portion of fluid on the other side of the barrier, with larger particles removed filtered through x posteriorly_connected_to y iff the posterior part of x is connected to y. i.e. x connected_to y and x anterior_to y. uberon posteriorly_connected_to posteriorly connected to x posteriorly_connected_to y iff the posterior part of x is connected to y. i.e. x connected_to y and x anterior_to y. uberon protects protects uberon subdivision_of placeholder relation. X = 'subdivision of A' and subdivision_of some B means that X is the mereological sum of A and B subdivision of . uberon transitively_anteriorly_connected_to transitively anteriorly connected to . uberon transitively_connected_to transitively_connected to A protein coding gene resilin in fruit fly. gene FlyBase:FBgn0034157 Category=external. resilin (fruit fly) A protein coding gene resilin in fruit fly. PRO:DNx The projection of the inner surface of the cuticle. cuticular internal depression imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The projection of the inner surface of the cuticle. 2021-05-19 jgiron https://orcid.org/0000-0002-0851-6883 The region of the cuticle that is composed two articular surfaces that are adjacent to each other. Articulations are `anatomical clusters`, collections of not direclty connected (continuous) sclerite surfaces that are adjacent to another sclerite. The cranio-mandibular articulation is the articular surface of the cranium and the adjacent articular surface of the mandible. The articular surfaces are sometimes located on the same sclerite. insect articulation imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of the cuticle that is composed two articular surfaces that are adjacent to each other. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 Articulations are `anatomical clusters`, collections of not direclty connected (continuous) sclerite surfaces that are adjacent to another sclerite. The cranio-mandibular articulation is the articular surface of the cranium and the adjacent articular surface of the mandible. The articular surfaces are sometimes located on the same sclerite. imiko https://orcid.org/0000-0003-2938-9075 The region of the insect cuticle that is less flexible than the neighboring conjunctiva(e) (conjunctiva(e) that the sclerite is continuous with). sclerite imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of the insect cuticle that is less flexible than the neighboring conjunctiva(e) (conjunctiva(e) that the sclerite is continuous with). imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 sclerite Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Sclerite: distinctly sclerotized region of the integument, with thick layer of exocuticle." The region of the insect cuticle that is more flexible than the neighboring sclerite(s) (sclerite(s) that the conjunctiva is continuous with). Conjunctivae are more flexible and usually less thick than sclerties of the cuticle allowing the sclerites to move relative to each other. Micro-CT based methods cannot be used to accurately differentiate slcerites from conjunctiuvae. CLSM (the excitation wavelengths of sclerites and conjunctivae are different in response of 488 nm emission vawelength), histological sections or simple dissections need to be applied to define relative flexibility of the insect cuticle. If a conjunctiva entirelly encircles a sclerite, then the 'encircles' and 'encircled by' object properties are used to link the sclerite and the conjunctiva even in cases in which the encircled sclerites or conjunctivae are part of an evagination allowing to model spatial relationship between appendage segments: the pedicel is encircled by the scapal-pedicellar conjunctiva, which is encircled by the scape or the the coxa encircles the coxo-femoral conjunctiuva that encircels the femur. membrane conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of the insect cuticle that is more flexible than the neighboring sclerite(s) (sclerite(s) that the conjunctiva is continuous with). imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 Conjunctivae are more flexible and usually less thick than sclerties of the cuticle allowing the sclerites to move relative to each other. Micro-CT based methods cannot be used to accurately differentiate slcerites from conjunctiuvae. CLSM (the excitation wavelengths of sclerites and conjunctivae are different in response of 488 nm emission vawelength), histological sections or simple dissections need to be applied to define relative flexibility of the insect cuticle. If a conjunctiva entirelly encircles a sclerite, then the 'encircles' and 'encircled by' object properties are used to link the sclerite and the conjunctiva even in cases in which the encircled sclerites or conjunctivae are part of an evagination allowing to model spatial relationship between appendage segments: the pedicel is encircled by the scapal-pedicellar conjunctiva, which is encircled by the scape or the the coxa encircles the coxo-femoral conjunctiuva that encircels the femur. imiko https://orcid.org/0000-0003-2938-9075 membrane Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Membrane: very flexible, unsclerotized area of the integument." The region of the cuticle that corresponds to a concave surface. External cuticular depression. The surface of the cuticle in insects is the entire surface of an organism, and it is composed of flat, concave and convex regions. Concavities sometimes correspons with invaginations of the cuticle and the underlaying sinlge layer epidermis, somtimes correspond to a decreased cuticular thickness. cuticular depression jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of the cuticle that corresponds to a concave surface. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 External cuticular depression. The surface of the cuticle in insects is the entire surface of an organism, and it is composed of flat, concave and convex regions. Concavities sometimes correspons with invaginations of the cuticle and the underlaying sinlge layer epidermis, somtimes correspond to a decreased cuticular thickness. imiko https://orcid.org/0000-0003-2938-9075 The region of cuticle that corresponds with an invagination of the single layer epidermis (epidermal fold). The cuticular invagination sometimes corresponds to a cuticular depression (concavity on the surface of the cuticle). cuticular invagination imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of cuticle that corresponds with an invagination of the single layer epidermis (epidermal fold). The cuticular invagination sometimes corresponds to a cuticular depression (concavity on the surface of the cuticle). imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The region of the cuticle with higher content of resilin. elastic cuticle imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of the cuticle with higher content of resilin. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 elastic cuticle 2020-10-27 The region of the cuticle that corresponds to a convex surface. The surface of the cuticle in insects is the entire surface of an organism, and it is composed of flat, concave and convex regions. Convexities sometimes correspons with evaginations of the cuticle and the underlaying sinlge layer epidermis and sometimes correspond to an increased cuticular thickness." cuticular protrusion jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 starasov https://orcid.org/0000-0001-5237-2330 imiko https://orcid.org/0000-0003-2938-9075 The region of the cuticle that corresponds to a convex surface. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The surface of the cuticle in insects is the entire surface of an organism, and it is composed of flat, concave and convex regions. Convexities sometimes correspons with evaginations of the cuticle and the underlaying sinlge layer epidermis and sometimes correspond to an increased cuticular thickness." imiko https://orcid.org/0000-0003-2938-9075 A part or structure that is produced by evagination. evaginated imiko https://orcid.org/0000-0003-2938-9075 A part or structure that is produced by evagination. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 evaginated imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The articulation between the mandible and the cranium. cranio-mandibular articulation imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The articulation between the mandible and the cranium. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The articulation between the cardo and the cranium. cranio-cardinal articulation imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The articulation between the cardo and the cranium. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The articulation between the scapus and the cranium. cranio-antennal articulation imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The articulation between the scapus and the cranium. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 The articulation between the head and the neck. head-cervical articulation imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The articulation between the head and the neck. imiko https://orcid.org/0000-0003-2938-9075 2020-10-27 Is the sclerite of the insect maxilla that is attached to the head capsule by the oral foraminal conjunctiva and to the stipes by the stipito-cardinal conjunctiva. cardo imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 Is the sclerite of the insect maxilla that is attached to the head capsule by the oral foraminal conjunctiva and to the stipes by the stipito-cardinal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 cardo Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Cardo: basal element of maxilla, articulating with head capsule; usually moved by one cranial muscle and one tentorial muscle." Is the sclerite of the insect head that is attached to the neck membrane and the oral foraminal conjunctiva, bears the eye cuticle and surrounds the brain insect cranium head capsule imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 Is the sclerite of the insect head that is attached to the neck membrane and the oral foraminal conjunctiva, bears the eye cuticle and surrounds the brain jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 insect cranium imiko https://orcid.org/0000-0003-2938-9075 head capsule Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "The head capsule is almost always a rigid, well sclerotized structural unit, equipped with a defined set of appendages. It is reinforced by strengthening ridges and the endoskeletal tentorium" The axillary sclerite that is on the dorsal region of the basal wing conjunctiva and lateral to the first axillary sclerite. dorsal second axillary sclerite imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The axillary sclerite that is on the dorsal region of the basal wing conjunctiva and lateral to the first axillary sclerite. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-10 Is the abdominal tergite of the preabdomen that is attached to the posterior margin of the tergum by the metanoto-abdominal tergite I conjunctiva and to the anterior margin of the abdominal tergite II by the abdominal tergite I-abdominal tergite II conjunctiva. abdominal tergite I jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the abdominal tergite of the preabdomen that is attached to the posterior margin of the tergum by the metanoto-abdominal tergite I conjunctiva and to the anterior margin of the abdominal tergite II by the abdominal tergite I-abdominal tergite II conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The axillary sclerite that is part of the dorsal region of the basal wing conjunctiva and is anterior to the third axillary sclerite. first axillary sclerite imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The axillary sclerite that is part of the dorsal region of the basal wing conjunctiva and is anterior to the third axillary sclerite. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-10 The appendage of the insect maxilla that is attached to the stipes by the stipito-galeal conjunctiva. galea jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The appendage of the insect maxilla that is attached to the stipes by the stipito-galeal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 galea Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Galea: outer endite lobe of maxilla, usually semimembranous and equipped with sensilla (mainly chemoreceptors)." The paired appendage of the labium that is attached to the palpiger by the palpigero-palpal conjunctiva. labial palpus imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The paired appendage of the labium that is attached to the palpiger by the palpigero-palpal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 labial palpus Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Labial palp: tactile appendage of the labium." The appendage of the insect maxilla that is attached to the stipes by the stipito-lacinial conjunctiva. lacinia jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The appendage of the insect maxilla that is attached to the stipes by the stipito-lacinial conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 lacinia Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Lacinia: inner endite lobe of maxilla, usually firmly connected or fused with stipes; usually sclerotized, with mesally curved apex and articulated spines arranged along mesal edge." The region of cuticle that corresponds with an evagination of the cuticle and the single layer epidermis (epidermal fold). The cuticular evagination usually corresponds to a cuticular protrusion (convexity on the surface of the cuticle). cuticular evagination jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The region of cuticle that corresponds with an evagination of the cuticle and the single layer epidermis (epidermal fold). The cuticular evagination usually corresponds to a cuticular protrusion (convexity on the surface of the cuticle). imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The cuticular protrusion that does not correspond to an evagination. solid cuticular protrusion imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The cuticular protrusion that does not correspond to an evagination. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 solid cuticular protrusion 2020-11-11 The evagination of the cuticle that is musculated (connected to the body via muscles). appendage jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The evagination of the cuticle that is musculated (connected to the body via muscles). imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 appendage Nichols, S. W. (1989) Torre-Bueno glossary of entomology. New York Entomological Society. 840 pp. https://ipmhouse.tamu.edu/files/2021/03/The-Torre-Bueno-Glossary-of-Entomology.pdf "any part, piece, or organ attached by a joint to the body or to any other main structure" Is the paired appendage of the insect thorax that is composed of coxa, trochanter, femur, tibia, tarsus, and pretarsus. insect leg jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the paired appendage of the insect thorax that is composed of coxa, trochanter, femur, tibia, tarsus, and pretarsus. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 Is the appendage of the head capsule that is connected by the antenniferal-scapal conjunctiva to the scapus, which is connected by the scapal-pedicellar conjunctiva to the pedicellus; the pedicellus is connected by the pedicellar-first flagellomeral conjunctiva to the first flagellomere; the first flagellomere is connected by interflagellomeral conjunctiva to the next flagellomere. Further flagellomeres are connected to each other by interflagellomeral conjunctiva. antenna imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the appendage of the head capsule that is connected by the antenniferal-scapal conjunctiva to the scapus, which is connected by the scapal-pedicellar conjunctiva to the pedicellus; the pedicellus is connected by the pedicellar-first flagellomeral conjunctiva to the first flagellomere; the first flagellomere is connected by interflagellomeral conjunctiva to the next flagellomere. Further flagellomeres are connected to each other by interflagellomeral conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 antenna Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Antenna: head appendage of segment 2, innervated by deutocerebrum; in Insecta composed of scapus, pedicellus and flagellum; primarily multisegmented and filiform." The flat, paired appendage on the dorso-lateral region of the pterothorax. insect wing imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The flat, paired appendage on the dorso-lateral region of the pterothorax. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The paired appendage of the prothorax that is anterior to a mid leg and composed of procoxa, protrochanter, profemur, protibia, protarsus, and propretarsus. fore leg jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The paired appendage of the prothorax that is anterior to a mid leg and composed of procoxa, protrochanter, profemur, protibia, protarsus, and propretarsus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The paired appendage of the mesothorax that is posterior to a fore leg, anterior to a hind leg, and composed of mesocoxa, mesotrochanter, mesofemur, mesotibia, mesotarsus, and mesopretarsus. mid leg jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The paired appendage of the mesothorax that is posterior to a fore leg, anterior to a hind leg, and composed of mesocoxa, mesotrochanter, mesofemur, mesotibia, mesotarsus, and mesopretarsus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The paired appendage of the metathorax that is posterior to a mid leg and composed of metacoxa, metatrochanter, metafemur, metatibia, metatarsus, and metapretarsus. hind leg jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The paired appendage of the metathorax that is posterior to a mid leg and composed of metacoxa, metatrochanter, metafemur, metatibia, metatarsus, and metapretarsus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The paired insect wing that is continuous with the mesonotum. fore wing imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The paired insect wing that is continuous with the mesonotum. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The paired insect wing that is continuous with the metanotum. hind wing imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The paired insect wing that is continuous with the metanotum. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The unicellular cuticular evagination that constitutes a sclerite with its base surrounded by socket membrane. seta imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The unicellular cuticular evagination that constitutes a sclerite with its base surrounded by socket membrane. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 seta Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Seta (=macrotrichium): articulated hair with socket, formed by trichogen cell." 2020-11-11 The multicellular evagination of the cuticle that corresponds to an epithelial fold, composed of some sclerite that is encricled by a conjunctiva (moveable). The difference between an appendage and a spur is the lack of muscles attached to the spur, otherwise, the mandible would be also a spur. Spines are multicellular evaginations that are not encircled by conjunctivae. Moveable beetle horns are spurs while not moveable beetle horns are spines. spur imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The multicellular evagination of the cuticle that corresponds to an epithelial fold, composed of some sclerite that is encricled by a conjunctiva (moveable). imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The difference between an appendage and a spur is the lack of muscles attached to the spur, otherwise, the mandible would be also a spur. Spines are multicellular evaginations that are not encircled by conjunctivae. Moveable beetle horns are spurs while not moveable beetle horns are spines. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 spur Richards AG, Richards PA (1979) The cuticular protuberances of insects. International Journal of Insect Morphology and Embryology 8: 143–157. "Multicellular protuberances without differentiation of specialized cells" https://doi.org/10.1016/0020-7322(79)90013-8 The seta that is flat. scale imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The seta that is flat. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 Is the mouthpart and appendage that is attached to the clypeus by the clypeo-labral conjunctiva and to the proximal margin of the epipharynx. labrum imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the mouthpart and appendage that is attached to the clypeus by the clypeo-labral conjunctiva and to the proximal margin of the epipharynx. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 labrum Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Labrum: unpaired upper lip, connected with distal edge of clypeus." Is a mouthpart and sclerite between the labrum and the insect maxilla that is attached to the head capsule by the oral foraminal conjunctiva and the cranio-mandibular muscle. insect mandible imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is a mouthpart and sclerite between the labrum and the insect maxilla that is attached to the head capsule by the oral foraminal conjunctiva and the cranio-mandibular muscle. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 insect mandible Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "The mandibles are the paired appendages of the 4th head segment and receive their innervation from the anterior part of the suboesophageal ganglion [...] They are primarily a compact, undivided and strongly sclerotized element without appendages." Is a mouthpart appendage of the head capsule, between the insect mandible and the labium, composed of cardo, which is attached to the stipes by the cardo-stipital conjunctiva; the stipes, which is attached to the galea by the stipito-galeal conjunctiva, to the lacinia by the stipito-lacinial conjunctiva, and to the palpifer by the stipito-palpiferal conjunctiva; the palpifer is attached to the maxillary palpus by the palpifero-palpal conjunctiva. insect maxilla imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is a mouthpart appendage of the head capsule, between the insect mandible and the labium, composed of cardo, which is attached to the stipes by the cardo-stipital conjunctiva; the stipes, which is attached to the galea by the stipito-galeal conjunctiva, to the lacinia by the stipito-lacinial conjunctiva, and to the palpifer by the stipito-palpiferal conjunctiva; the palpifer is attached to the maxillary palpus by the palpifero-palpal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 insect maxilla Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Maxilla: head appendage of segment 5, innervated by maxillary nerve originating from intermediate part of suboesophageal complex; primarily composed of cardo, stipes, endite lobes (lacinia, galea) and maxillary palp." The conjunctiva of the insect thorax that connects the mesothorax and the metathorax. mesothoracic-metathoracic conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The conjunctiva of the insect thorax that connects the mesothorax and the metathorax. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The conjunctiva of the insect thorax joining the prothorax and the mesothorax. prothoracic-mesothoracic conjunctiva jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect thorax joining the prothorax and the mesothorax. jgiron https://orcid.org/0000-0002-0851-6883 2020-03-18 The appendage of the leg that is attached to the tarsus by the tarso-pretarsal conjunctiva. ungues pretarsus jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The appendage of the leg that is attached to the tarsus by the tarso-pretarsal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 ungues Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Ungues: pretarsal claws; unpaired in Collembola and Protura, usually paired in Diplura and Insecta." Is the appendage of the labium composed of the glossa and paraglossa jointly. ligula imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the appendage of the labium composed of the glossa and paraglossa jointly. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 ligula Snodgrass RE (1993) Principles of Insect Morphology. Cornell University Press, 768 pp. https://www.jstor.org/stable/10.7591/j.ctv1nhm1j "Ligula - The terminal lobes of the labium collectively, or a terminal part of the labium formed by the union of the lobes." Is the appendage of the labium, and part of the ligula, that is attached to the prementum by the premental-glossal conjunctiva. glossa imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the appendage of the labium, and part of the ligula, that is attached to the prementum by the premental-glossal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 glossa Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Glossa: paired inner endite lobes of the prementum." The paired appendage of the labium, and part of the ligula, that is attached to the prementum by the premental-paraglossal conjunctiva. paraglossa imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The paired appendage of the labium, and part of the ligula, that is attached to the prementum by the premental-paraglossal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 paraglossa Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Paraglossa: paired outer endite lobes of the prementum (labium)." Is the appendage of the insect maxilla that is attached to the palpifer by the palpifero-palpal conjunctiva. maxillary palpus imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the appendage of the insect maxilla that is attached to the palpifer by the palpifero-palpal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 maxillary palpus Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Palp: tactile appendage of maxilla" The notum or sclerite on the dorsal region of the mesothorax that is posterior to the pronotum, connected via the mesonoto-mesopleural conjuntiva to the dorsal margin of the mesopleuron, that is not adjacent to the mesocoxa, but attached to it by the mesocoxal muscle. mesonotum jcgiron imiko https://orcid.org/0000-0003-2938-9075 jcgiron jgiron https://orcid.org/0000-0002-0851-6883 jcgiron jcgiron jcgiron jcgiron The notum or sclerite on the dorsal region of the mesothorax that is posterior to the pronotum, connected via the mesonoto-mesopleural conjuntiva to the dorsal margin of the mesopleuron, that is not adjacent to the mesocoxa, but attached to it by the mesocoxal muscle. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The sclerite that is continuous with the basal mesocoxal conjunctiva and the basal wing conjunctiva and contains the mesofurca. mesopectus imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The sclerite that is continuous with the basal mesocoxal conjunctiva and the basal wing conjunctiva and contains the mesofurca. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 mesopectus 2020-11-11 The trochantin that is located on the ventro-lateral region of the mesothorax and is attached to a mesocoxa and surrounded by the basal mesocoxal conjunctiva. mesotrochantin imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The trochantin that is located on the ventro-lateral region of the mesothorax and is attached to a mesocoxa and surrounded by the basal mesocoxal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 mesotrochantin 2020-11-11 The trochantin that is located on the ventro-lateral region of the prothorax and is attached to a procoxa and surrounded by the basal procoxal conjunctiva. protrochantin imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The trochantin that is located on the ventro-lateral region of the prothorax and is attached to a procoxa and surrounded by the basal procoxal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 protrochantin 2020-11-11 The trochantin that is located on the ventro-lateral region of the metathorax and is attached to a metacoxa and surrounded by the basal metacoxal conjunctiva. metatrochantin imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The trochantin that is located on the ventro-lateral region of the metathorax and is attached to a metacoxa and surrounded by the basal metacoxal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 metatrochantin 2020-11-11 Is the sclerite of the labium that is connected to the mentum by the mental-premental conjunctiva, to the palpiger by the premental-palpigeral conjunctiva, to the paraglossa by the premental-paraglossal conjunctiva, and to the glossa by the premental-glossal conjunctiva. prementum imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the sclerite of the labium that is connected to the mentum by the mental-premental conjunctiva, to the palpiger by the premental-palpigeral conjunctiva, to the paraglossa by the premental-paraglossal conjunctiva, and to the glossa by the premental-glossal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 prementum Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Prementum: distal main element of labium, bearing the endite lobes (glossa and paraglossa) and the labial palp." The sclerite that is posterior to the pronotum and receives the site of attachment of the dilutor muscle of the first thoracic spiracle. prepectus imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The sclerite that is posterior to the pronotum and receives the site of attachment of the dilutor muscle of the first thoracic spiracle. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 prepectus 2020-11-11 The sclerite that is attached to cranial muscles, procoxal muscles, contiuous posterioirly with the neck membrane and with pronoto-propleural conjunctiva and not adjacent (not touching) to the procoxa. pronotum imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The sclerite that is attached to cranial muscles, procoxal muscles, contiuous posterioirly with the neck membrane and with pronoto-propleural conjunctiva and not adjacent (not touching) to the procoxa. imiko https://orcid.org/0000-0003-2938-9075 2021-03-18 pronotum Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Pronotum: prothoracic tergum." The mere that is part of the labial palpus. labial palpomere jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The mere that is part of the labial palpus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The sclerite on the ventral region of the prothorax that bears the profurca and us attached to the propleuro-prosternal conjunctiva. prosternum imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The sclerite on the ventral region of the prothorax that bears the profurca and us attached to the propleuro-prosternal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The sclerite that is cylindrical and encircles and is encircled by conjunctiva. ring sclerite imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The sclerite that is cylindrical and encircles and is encircled by conjunctiva. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 The ring sclerite that is part of an appendage and is attached to muscle. appendage segment imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The ring sclerite that is part of an appendage and is attached to muscle. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 The segment of the mid leg that is attached to the latero-ventral region of the mesothorax by the basal mesocoxal conjunctiva and to the mesotrochanter by the mesocoxal-mesotrochanteral conjunctiva. mesocoxa imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of the mid leg that is attached to the latero-ventral region of the mesothorax by the basal mesocoxal conjunctiva and to the mesotrochanter by the mesocoxal-mesotrochanteral conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The segment of the hind leg that is attached to the latero-ventral region of the metathorax by the basal metacoxal conjunctiva and to the metatrochanter by the metacoxal-metatrochanteral conjunctiva. metacoxa imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of the hind leg that is attached to the latero-ventral region of the metathorax by the basal metacoxal conjunctiva and to the metatrochanter by the metacoxal-metatrochanteral conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The segment of the fore leg that is attached to the latero-ventral region of the prothorax by the basal procoxal conjunctiva and to the protrochanter by the procoxal-protrochanteral conjunctiva. procoxa imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of the fore leg that is attached to the latero-ventral region of the prothorax by the basal procoxal conjunctiva and to the protrochanter by the procoxal-protrochanteral conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The segment of a fore leg attached to the distal margin of the profemur by the profemoro-protibial conjunctiva and to the proximal margin of the protarsus by the protibio-protarsal conjunctiva. protibia jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a fore leg attached to the distal margin of the profemur by the profemoro-protibial conjunctiva and to the proximal margin of the protarsus by the protibio-protarsal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The segment of a mid leg attached to the distal margin of the mesofemur by the mesofemoro-mesotibial conjunctiva and to the proximal margin of the mesotarsus by the mesotibio-mesotarsal conjunctiva. mesotibia jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a mid leg attached to the distal margin of the mesofemur by the mesofemoro-mesotibial conjunctiva and to the proximal margin of the mesotarsus by the mesotibio-mesotarsal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The segment of a hind leg attached to the distal margin of the metafemur by the metafemoro-metatibial conjunctiva and to the proximal margin of the metatarsus by the metatibio-metatarsal conjunctiva. metatibia jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a hind leg attached to the distal margin of the metafemur by the metafemoro-metatibial conjunctiva and to the proximal margin of the metatarsus by the metatibio-metatarsal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The segment of a fore leg attached to the distal margin of the protrochanter by the protrochantero-profemoral conjunctiva and to the proximal margin of the protibia by the profemoro-protibial conjunctiva. profemur jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a fore leg attached to the distal margin of the protrochanter by the protrochantero-profemoral conjunctiva and to the proximal margin of the protibia by the profemoro-protibial conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The segment of a mid leg attached to the distal margin of the mesotrochanter by the mesotrochantero-mesofemoral conjunctiva and to the proximal margin of the mesotibia by the mesofemoro-mesotibial conjunctiva. mesofemur jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a mid leg attached to the distal margin of the mesotrochanter by the mesotrochantero-mesofemoral conjunctiva and to the proximal margin of the mesotibia by the mesofemoro-mesotibial conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The segment of a hind leg attached to the distal margin of the metatrochanter by the metatrochantero-metafemoral conjunctiva and to the proximal margin of the metatibia by the metafemoro-metatibial conjunctiva. metafemur jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a hind leg attached to the distal margin of the metatrochanter by the metatrochantero-metafemoral conjunctiva and to the proximal margin of the metatibia by the metafemoro-metatibial conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The ring sclerite that is part of an appendage and is not attached to skeletal muscle tissue. mere imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The ring sclerite that is part of an appendage and is not attached to skeletal muscle tissue. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The mere that is part of the tarsus. tarsomere imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The mere that is part of the tarsus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 tarsomere "Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Tarsomere: subelement of tarsus in Insecta; number often taxon-specific, sometimes differing on legs of different segments (diagnostic value: tarsal formula)." The segment of a leg attached to the distal margin of the femur by the femoro-tibial conjunctiva and to the proximal margin of the tarsus by the tibio-tarsal conjunctiva. tibia jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a leg attached to the distal margin of the femur by the femoro-tibial conjunctiva and to the proximal margin of the tarsus by the tibio-tarsal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 tibia Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Tibia: 4th element of leg; usually cylindrical and equipped with paired terminal spurs, often with rows of spines." The segment of a leg attached to the distal margin of the trochanter by the trochantero-femoral conjunctiva and to the proximal margin of the tibia by the femoro-tibial conjunctiva. femur jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of a leg attached to the distal margin of the trochanter by the trochantero-femoral conjunctiva and to the proximal margin of the tibia by the femoro-tibial conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 femur Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Femur: 3rd element of leg, often largest part." The segment of the leg that is attached to the latero-ventral region of the insect thorax by the basal coxal conjunctiva and to the trochanter by the coxal-trochanteral conjunctiva. coxa imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of the leg that is attached to the latero-ventral region of the insect thorax by the basal coxal conjunctiva and to the trochanter by the coxal-trochanteral conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 coxa Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Coxa: basal element of leg; articulated with pleural, sternal and usually also trochantinal elements of thoracic segment." The skeletal muscle continuous with the head capsule. insect cranial muscle The skeletal muscle continuous with the head capsule. lagonzalezmo https://orcid.org/0000-0002-9136-9932 2021-04-04 The skeletal muscle continuous with the head capsule and the antenna. cranio-antennal muscle lagonzalezmo https://orcid.org/0000-0002-9136-9932 lagonzalezmo https://orcid.org/0000-0002-9136-9932 The skeletal muscle continuous with the head capsule and the antenna. lagonzalezmo https://orcid.org/0000-0002-9136-9932 2021-04-04 The skeletal muscle continuous with the head capsule and the insect mandible. cranio-mandibular muscle imiko https://orcid.org/0000-0003-2938-9075 lagonzalezmo https://orcid.org/0000-0002-9136-9932 The skeletal muscle continuous with the head capsule and the insect mandible. lagonzalezmo https://orcid.org/0000-0002-9136-9932 2021-04-04 The skeletal muscle continuous with the head capsule and the labium. cranio-labial muscle lagonzalezmo https://orcid.org/0000-0002-9136-9932 lagonzalezmo https://orcid.org/0000-0002-9136-9932 The skeletal muscle continuous with the head capsule and the labium. lagonzalezmo https://orcid.org/0000-0002-9136-9932 2021-04-04 The skeletal muscle continuous with the head capsule and the labrum. cranio-labral muscle lagonzalezmo https://orcid.org/0000-0002-9136-9932 lagonzalezmo https://orcid.org/0000-0002-9136-9932 The skeletal muscle continuous with the head capsule and the labrum. lagonzalezmo https://orcid.org/0000-0002-9136-9932 2021-04-04 The coxal muscle of the mesocoxa. mesocoxal muscle imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The coxal muscle of the mesocoxa. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 mesocoxal muscle 2020-11-11 The skeletal muscle that inserts in the coxa. All of the coxal muscles, except the second sternal-metacoxal muscle, are attached to the thorax. coxal muscle imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The skeletal muscle that inserts in the coxa. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 All of the coxal muscles, except the second sternal-metacoxal muscle, are attached to the thorax. imiko https://orcid.org/0000-0003-2938-9075 Is the mouthpart appendage composed of the submentum, which is attached by the submental-mental conjunctiva to the mentum, which is attached by the mental-premental conjunctiva to the prementum, which is attached to the palpiger by the premental-palpigeral conjunctiva, the paraglossa by the premental-paraglossal conjunctiva, and the glossa by the premental-glossal conjunctiva; the palpiger is attached by the palpigero-palpal conjunctiva to the labial palpus. labium imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the mouthpart appendage composed of the submentum, which is attached by the submental-mental conjunctiva to the mentum, which is attached by the mental-premental conjunctiva to the prementum, which is attached to the palpiger by the premental-palpigeral conjunctiva, the paraglossa by the premental-paraglossal conjunctiva, and the glossa by the premental-glossal conjunctiva; the palpiger is attached by the palpigero-palpal conjunctiva to the labial palpus. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 labium Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "The labium or lower lip forms the posterior (orthognathous head) or ventral (prognathous head) closure of the preoral space and of the salivarium. It is homologous to the maxilla (2nd maxilla) of crustaceans, but its proximal parts form a structural unit without a recognizable median line or suture." The elastic cuticle of the head capsule that is transparent and covers the eye. eye cuticle imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The elastic cuticle of the head capsule that is transparent and covers the eye. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 eye cuticle 2020-11-11 The resilin rich region that connects two sclerites. ligament imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The resilin rich region that connects two sclerites. imiko https://orcid.org/0000-0003-2938-9075 2021-04-21 ligament 2020-11-11 The usually cylindircal evagination of the cuticle that is resilin rich and is continuous with skeletal muscles. cuticular tendon imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The usually cylindircal evagination of the cuticle that is resilin rich and is continuous with skeletal muscles. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 cuticular tendon Klass K-D (2001) The female abdomen of the viviparous earwig Hemimerus vosseleri (Insecta: Dermaptera: Hemimeridae), with a discussion of the postgenital abdomen of Insecta. Zoological Journal of the Linnean Society 131: 251–307. https://doi.org/10.1111/j.1096-3642.2001.tb02239.x "tendon is used herein to designate membranous internal cuticular projections" The branching invagination of elastic cuticle that is cylindrical and contains taenidia. trachea imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The branching invagination of elastic cuticle that is cylindrical and contains taenidia. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 trachea Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Trachea: tube-shaped elements of the tracheal system." The conjunctiva of the insect head that is attached to the salivarium and to the oral foraminal conjunctiva. hypopharynx imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect head that is attached to the salivarium and to the oral foraminal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 hypopharynx Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Hypopharynx: median unpaired lobe between mouthparts and epipharynx; anterior (dorsal) surface continuous with anterior pharynx." The conjunctiva of the insect thorax that joins the mesopleuron and the metapleuron. mesopleuro-metapleural conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect thorax that joins the mesopleuron and the metapleuron. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjunctiva of the mesothorax that joins the metasternum and the metacoxa. metasterno-metacoxal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the mesothorax that joins the metasternum and the metacoxa. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjunctiva of the prothorax that joins the prosternum and the procoxa. prosternal-procoxal conjunctiva jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the prothorax that joins the prosternum and the procoxa. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjunctiva of the metathorax that joins the metapleuron and the metacoxa. metapleuro-metacoxal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the metathorax that joins the metapleuron and the metacoxa. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjuctiva that is continuous with the posterior margin of the head capsule and with the anterior margin of the prnotum, and bears cervical sclerites. cervical membrane neck membrane imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The conjuctiva that is continuous with the posterior margin of the head capsule and with the anterior margin of the prnotum, and bears cervical sclerites. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 cervical membrane Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Cervical membrane: membrane connecting the head and prothorax." The conjunctiva of the insect head that connects the head capsule and the mouthparts. oral foraminal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect head that connects the head capsule and the mouthparts. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The conjunctiva of the insect thorax that joins the pronotum and the mesonotum. pronoto-mesonotal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect thorax that joins the pronotum and the mesonotum. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjunctiva that is attached to the pronotum and the mesopectus. pronoto-mesopectal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva that is attached to the pronotum and the mesopectus. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The conjunctiva that encircles a ring sclerite. ring conjunctiva imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The conjunctiva that encircles a ring sclerite. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-03 The conjunctiva that encircles the mesocoxa and connects it to the ventral and lateral regions of the mesothorax. basal mesocoxal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva that encircles the mesocoxa and connects it to the ventral and lateral regions of the mesothorax. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The conjunctiva that encircles the metacoxa and connects it to the ventral and lateral regions of the metathorax. basal metacoxal conjunctiva jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva that encircles the metacoxa and connects it to the ventral and lateral regions of the metathorax. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The conjunctiva of the insect head that is connected to the oral foraminal conjunctiva and to the distal margin of the labrum. epipharynx imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The conjunctiva of the insect head that is connected to the oral foraminal conjunctiva and to the distal margin of the labrum. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 epipharynx Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Epipharynx: semimembranous or membranous posterior wall of labrum and roof of cibarium." The conjunctiva that encircles the procoxa and connects it to the ventral and lateral regions of the prothorax. basal procoxal conjunctiva jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva that encircles the procoxa and connects it to the ventral and lateral regions of the prothorax. jgiron https://orcid.org/0000-0002-0851-6883 2021-03-18 The cuticular region that is anterior (distal) to the neck membrane (encircled by the neck membrane) and contains the head capsule, the antenna, the eyes, mouthparts and surrounds the brain. insect head jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The cuticular region that is anterior (distal) to the neck membrane (encircled by the neck membrane) and contains the head capsule, the antenna, the eyes, mouthparts and surrounds the brain. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 insect head Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "The head is a compact and complex tagma with a concentration of a broad array of structures und functions. It is equipped with sense organs such as the highly complex compound eyes and the antennae, an elaborate set of mouthparts, and a complex muscle apparatus. It also contains central elements of the nervous system, the anterior part of the digestive tract including the cibarium (preoral cavity) and the salivarium, and also neurohaemal organs." The region of cuticle that is connected to some muscles of the fore and hind legs and is posterior to the neck membrane and aterior to the first abdominal spiracle. The hind leg muscles sometimes attach to the second abdominal sternite. The abdominal sopiracles are different in their structure from tghe thoracic spiracles as they are opened and closed by two articulating sclerites that are moved by two sets of muscles whereas the thoracic spiracles are closed by only one occlusor muscle and opened by the flexibility of the distal tracheal region. The border between the thorax and the abdomen is sometimes marked by the anterior first abdmonial tergal conjunctiva. In Hymenoptera, the border between the abdomen and the thorax cannot be defined as the anterior first abdmonial tergal conjunctiva is absent, the sclerite that receives muscles from the metacoxa also contains the first abdominal spiracle. insect thorax jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The region of cuticle that is connected to some muscles of the fore and hind legs and is posterior to the neck membrane and aterior to the first abdominal spiracle. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The hind leg muscles sometimes attach to the second abdominal sternite. The abdominal sopiracles are different in their structure from tghe thoracic spiracles as they are opened and closed by two articulating sclerites that are moved by two sets of muscles whereas the thoracic spiracles are closed by only one occlusor muscle and opened by the flexibility of the distal tracheal region. The border between the thorax and the abdomen is sometimes marked by the anterior first abdmonial tergal conjunctiva. In Hymenoptera, the border between the abdomen and the thorax cannot be defined as the anterior first abdmonial tergal conjunctiva is absent, the sclerite that receives muscles from the metacoxa also contains the first abdominal spiracle. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The region of the cuticle that is posterior to the thorax. The thorax is defined based on local properties (e.g. attachment sites of coxal muscles) and therefore we can reference this class for the definition of the abdomen. insect abdomen jgiron https://orcid.org/0000-0002-0851-6883 2021-08-01 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 The region of the cuticle that is posterior to the thorax. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The thorax is defined based on local properties (e.g. attachment sites of coxal muscles) and therefore we can reference this class for the definition of the abdomen. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The conjunctiva of the insect maxilla that connects the cardo and the stipes. cardo-stipital conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the insect maxilla that connects the cardo and the stipes. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 Is the sclerite of the insect maxilla that is attached to the cardo by the cardo-stipital conjunctiva, attached to the lacinia by the stipito-lacinial conjunctiva, attached to the palpifer by the stipito-palpiferal conjunctiva, and attached to the galea by the stipito-galeal conjunctiva. stipes imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the sclerite of the insect maxilla that is attached to the cardo by the cardo-stipital conjunctiva, attached to the lacinia by the stipito-lacinial conjunctiva, attached to the palpifer by the stipito-palpiferal conjunctiva, and attached to the galea by the stipito-galeal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 stipes Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Stipes: second element of maxilla, connected with cardo by a hinge; bearing endite lobes (galea and lacinia) and maxillary palp." The conjunctiva that encircles the base of the antenna (scape) and is encircled by the head capsule as the antennal foramen. cranio-antennal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The conjunctiva that encircles the base of the antenna (scape) and is encircled by the head capsule as the antennal foramen. imiko https://orcid.org/0000-0003-2938-9075 2020-11-11 The segment of the antenna that is connected to the head capsule and the antennifer by the basal antennal conjunctiva and to the pedicellus by the scapal-pedicellar conjunctiva. scapus imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The segment of the antenna that is connected to the head capsule and the antennifer by the basal antennal conjunctiva and to the pedicellus by the scapal-pedicellar conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 scapus Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 Scapus: basal element of antenna, with two muscles attached to the pedicellus. The conjunctiva of the antenna that connects the scapus and the pedicellus. scapal-pedicellar conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the antenna that connects the scapus and the pedicellus. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 Is the segment of the antenna that is attached to the scapus by the scapal-pedicellar conjunctiva and to the first flagellomere by the pedicel-first flagellomere conjunctiva. pedicellus imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 Is the segment of the antenna that is attached to the scapus by the scapal-pedicellar conjunctiva and to the first flagellomere by the pedicel-first flagellomere conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 pedicellus Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Pedicellus: second antennal segment in Insecta; contains Johnston’s organ and lacks intrinsic muscles." The mere that is part of the antenna. flagellomere imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 The mere that is part of the antenna. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The first flagellomere of the antenna, attached to the pedicel by the pedicel-first flagellomere conjunctiva. first flagellomere jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The first flagellomere of the antenna, attached to the pedicel by the pedicel-first flagellomere conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 first flagellomere Beutel RG, Friedrich F, Yang X-K, Ge S-Q (2014) Insect Morphology and Phylogeny Insect morphology and phylogeny: a textbook for students of entomology. De Gruyter. https://doi.org/10.1515/9783110264043 "Flagellomere: segment of the flagellum." The conjunctiva of the antenna that connects the pedicellus and the first flagellomere. pedicellar-first flagellomeral conjunctiva imiko https://orcid.org/0000-0003-2938-9075 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the antenna that connects the pedicellus and the first flagellomere. jgiron https://orcid.org/0000-0002-0851-6883 2021-02-19 The region of the cuticle on the lateral region of the metathorax that is attached to the metanotum and the metasternum, and is attached to the metacoxa by the metacoxal muscle and the basal metacoxal conjunctiva. metapleuron imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The region of the cuticle on the lateral region of the metathorax that is attached to the metanotum and the metasternum, and is attached to the metacoxa by the metacoxal muscle and the basal metacoxal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-21 The conjunctiva of the prothorax that joins the propleuron and the procoxa. propleuro-procoxal conjunctiva imiko https://orcid.org/0000-0003-2938-9075 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The conjunctiva of the prothorax that joins the propleuron and the procoxa. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The sclerite on the ventral region of the mesothorax that bears the mesofurca and is attached to the mesopleuro-mesosternal conjunctiva. mesosternum jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883 The sclerite on the ventral region of the mesothorax that bears the mesofurca and is attached to the mesopleuro-mesosternal conjunctiva. jgiron https://orcid.org/0000-0002-0851-6883 2021-04-09 The conjunctiva of the fore leg joining the distal margin of a procoxa and the proximal margin of a protrochanter. procoxal-protrochanteral conjunctiva jgiron https://orcid.org/0000-0002-0851-6883 jgiron https://orcid.org/0000-0002-0851-6883