Ontology about the gross anatomy of the C. elegans C. elegans Gross Anatomy Ontology http://creativecommons.org/licenses/by/4.0/ OBO-Edit 2.1-beta16 19:10:2018 10:30 worm_anatomy 1.2 raymond $ definition term replaced by If R <- P o Q is a defining property chain axiom, then it also holds that R -> P o Q. Note that this cannot be expressed directly in OWL is a defining property chain axiom If R <- P o Q is a defining property chain axiom, then (1) R -> P o Q holds and (2) Q is either reflexive or locally reflexive. A corollary of this is that P SubPropertyOf R. is a defining property chain axiom where second argument is reflexive has_alternative_id database_cross_reference has_exact_synonym has_narrow_synonym has_obo_format_version has_obo_namespace has_related_synonym shorthand is part of my brain is part of my body (continuant parthood, two material entities) my stomach cavity is part of my stomach (continuant parthood, immaterial entity is part of material entity) this day is part of this year (occurrent parthood) a core relation that holds between a part and its whole Everything is part of itself. Any part of any part of a thing is itself part of that thing. Two distinct things cannot be part of each other. Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ Parthood requires the part and the whole to have compatible classes: only an occurrent can be part of an occurrent; only a process can be part of a process; only a continuant can be part of a continuant; only an independent continuant can be part of an independent continuant; only an immaterial entity can be part of an immaterial entity; only a specifically dependent continuant can be part of a specifically dependent continuant; only a generically dependent continuant can be part of a generically dependent continuant. (This list is not exhaustive.) A continuant cannot be part of an occurrent: use 'participates in'. An occurrent cannot be part of a continuant: use 'has participant'. A material entity cannot be part of an immaterial entity: use 'has location'. A specifically dependent continuant cannot be part of an independent continuant: use 'inheres in'. An independent continuant cannot be part of a specifically dependent continuant: use 'bearer of'. part_of BFO:0000050 worm_anatomy part_of part_of part of part_of http://www.obofoundry.org/ro/#OBO_REL:part_of has part my body has part my brain (continuant parthood, two material entities) my stomach has part my stomach cavity (continuant parthood, material entity has part immaterial entity) this year has part this day (occurrent parthood) a core relation that holds between a whole and its part Everything has itself as a part. Any part of any part of a thing is itself part of that thing. Two distinct things cannot have each other as a part. Occurrents are not subject to change and so parthood between occurrents holds for all the times that the part exists. Many continuants are subject to change, so parthood between continuants will only hold at certain times, but this is difficult to specify in OWL. See http://purl.obolibrary.org/obo/ro/docs/temporal-semantics/ Parthood requires the part and the whole to have compatible classes: only an occurrent have an occurrent as part; only a process can have a process as part; only a continuant can have a continuant as part; only an independent continuant can have an independent continuant as part; only a specifically dependent continuant can have a specifically dependent continuant as part; only a generically dependent continuant can have a generically dependent continuant as part. (This list is not exhaustive.) A continuant cannot have an occurrent as part: use 'participates in'. An occurrent cannot have a continuant as part: use 'has participant'. An immaterial entity cannot have a material entity as part: use 'location of'. An independent continuant cannot have a specifically dependent continuant as part: use 'bearer of'. A specifically dependent continuant cannot have an independent continuant as part: use 'inheres in'. has_part has part preceded by x is preceded by y if and only if the time point at which y ends is before or equivalent to the time point at which x starts. Formally: x preceded by y iff ω(y) <= α(x), where α is a function that maps a process to a start point, and ω is a function that maps a process to an end point. An example is: translation preceded_by transcription; aging preceded_by development (not however death preceded_by aging). Where derives_from links classes of continuants, preceded_by links classes of processes. Clearly, however, these two relations are not independent of each other. Thus if cells of type C1 derive_from cells of type C, then any cell division involving an instance of C1 in a given lineage is preceded_by cellular processes involving an instance of C. The assertion P preceded_by P1 tells us something about Ps in general: that is, it tells us something about what happened earlier, given what we know about what happened later. Thus it does not provide information pointing in the opposite direction, concerning instances of P1 in general; that is, that each is such as to be succeeded by some instance of P. Note that an assertion to the effect that P preceded_by P1 is rather weak; it tells us little about the relations between the underlying instances in virtue of which the preceded_by relation obtains. Typically we will be interested in stronger relations, for example in the relation immediately_preceded_by, or in relations which combine preceded_by with a condition to the effect that the corresponding instances of P and P1 share participants, or that their participants are connected by relations of derivation, or (as a first step along the road to a treatment of causality) that the one process in some way affects (for example, initiates or regulates) the other. is preceded by preceded_by http://www.obofoundry.org/ro/#OBO_REL:preceded_by preceded by precedes x precedes y if and only if the time point at which x ends is before or equivalent to the time point at which y starts. Formally: x precedes y iff ω(x) <= α(y), where α is a function that maps a process to a start point, and ω is a function that maps a process to an end point. precedes inheres in this fragility is a characteristic of this vase this red color is a characteristic of this apple a relation between a specifically dependent continuant (the characteristic) and any other entity (the bearer), in which the characteristic depends on the bearer for its existence. inheres_in Note that this relation was previously called "inheres in", but was changed to be called "characteristic of" because BFO2 uses "inheres in" in a more restricted fashion. This relation differs from BFO2:inheres_in in two respects: (1) it does not impose a range constraint, and thus it allows qualities of processes, as well as of information entities, whereas BFO2 restricts inheres_in to only apply to independent continuants (2) it is declared functional, i.e. something can only be a characteristic of one thing. characteristic of bearer of this apple is bearer of this red color this vase is bearer of this fragility Inverse of characteristic_of A bearer can have many dependents, and its dependents can exist for different periods of time, but none of its dependents can exist when the bearer does not exist. bearer_of is bearer of has characteristic participates in this blood clot participates in this blood coagulation this input material (or this output material) participates in this process this investigator participates in this investigation a relation between a continuant and a process, in which the continuant is somehow involved in the process participates_in participates in has participant this blood coagulation has participant this blood clot this investigation has participant this investigator this process has participant this input material (or this output material) a relation between a process and a continuant, in which the continuant is somehow involved in the process Has_participant is a primitive instance-level relation between a process, a continuant, and a time at which the continuant participates in some way in the process. The relation obtains, for example, when this particular process of oxygen exchange across this particular alveolar membrane has_participant this particular sample of hemoglobin at this particular time. has_participant http://www.obofoundry.org/ro/#OBO_REL:has_participant has participant this catalysis function is a function of this enzyme a relation between a function and an independent continuant (the bearer), in which the function specifically depends on the bearer for its existence A function inheres in its bearer at all times for which the function exists, however the function need not be realized at all the times that the function exists. function_of is function of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. function of this red color is a quality of this apple a relation between a quality and an independent continuant (the bearer), in which the quality specifically depends on the bearer for its existence A quality inheres in its bearer at all times for which the quality exists. is quality of quality_of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. quality of this investigator role is a role of this person a relation between a role and an independent continuant (the bearer), in which the role specifically depends on the bearer for its existence A role inheres in its bearer at all times for which the role exists, however the role need not be realized at all the times that the role exists. is role of role_of This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. role of this enzyme has function this catalysis function (more colloquially: this enzyme has this catalysis function) a relation between an independent continuant (the bearer) and a function, in which the function specifically depends on the bearer for its existence A bearer can have many functions, and its functions can exist for different periods of time, but none of its functions can exist when the bearer does not exist. A function need not be realized at all the times that the function exists. has_function has function this apple has quality this red color a relation between an independent continuant (the bearer) and a quality, in which the quality specifically depends on the bearer for its existence A bearer can have many qualities, and its qualities can exist for different periods of time, but none of its qualities can exist when the bearer does not exist. has_quality has quality this person has role this investigator role (more colloquially: this person has this role of investigator) a relation between an independent continuant (the bearer) and a role, in which the role specifically depends on the bearer for its existence A bearer can have many roles, and its roles can exist for different periods of time, but none of its roles can exist when the bearer does not exist. A role need not be realized at all the times that the role exists. has_role has role a relation between an independent continuant (the bearer) and a disposition, in which the disposition specifically depends on the bearer for its existence has disposition inverse of has disposition This relation is modeled after the BFO relation of the same name which was in BFO2, but is used in a more restricted sense - specifically, we model this relation as functional (inherited from characteristic-of). Note that this relation is now removed from BFO2020. disposition of A 'has regulatory component activity' B if A and B are GO molecular functions (GO_0003674), A has_component B and A is regulated by B. dos 2017-05-24T09:30:46Z has regulatory component activity A relationship that holds between a GO molecular function and a component of that molecular function that negatively regulates the activity of the whole. More formally, A 'has regulatory component activity' B iff :A and B are GO molecular functions (GO_0003674), A has_component B and A is negatively regulated by B. dos 2017-05-24T09:31:01Z By convention GO molecular functions are classified by their effector function. Internal regulatory functions are treated as components. For example, NMDA glutmate receptor activity is a cation channel activity with positive regulatory component 'glutamate binding' and negative regulatory components including 'zinc binding' and 'magnesium binding'. has negative regulatory component activity A relationship that holds between a GO molecular function and a component of that molecular function that positively regulates the activity of the whole. More formally, A 'has regulatory component activity' B iff :A and B are GO molecular functions (GO_0003674), A has_component B and A is positively regulated by B. dos 2017-05-24T09:31:17Z By convention GO molecular functions are classified by their effector function and internal regulatory functions are treated as components. So, for example calmodulin has a protein binding activity that has positive regulatory component activity calcium binding activity. Receptor tyrosine kinase activity is a tyrosine kinase activity that has positive regulatory component 'ligand binding'. has positive regulatory component activity dos 2017-05-24T09:44:33Z A 'has component activity' B if A is A and B are molecular functions (GO_0003674) and A has_component B. has component activity w 'has process component' p if p and w are processes, w 'has part' p and w is such that it can be directly disassembled into into n parts p, p2, p3, ..., pn, where these parts are of similar type. dos 2017-05-24T09:49:21Z has component process dos 2017-09-17T13:52:24Z Process(P2) is directly regulated by process(P1) iff: P1 regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding regulates the kinase activity (P2) of protein B then P1 directly regulates P2. directly regulated by Process(P2) is directly regulated by process(P1) iff: P1 regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding regulates the kinase activity (P2) of protein B then P1 directly regulates P2. GOC:dos Process(P2) is directly negatively regulated by process(P1) iff: P1 negatively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding negatively regulates the kinase activity (P2) of protein B then P2 directly negatively regulated by P1. dos 2017-09-17T13:52:38Z directly negatively regulated by Process(P2) is directly negatively regulated by process(P1) iff: P1 negatively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding negatively regulates the kinase activity (P2) of protein B then P2 directly negatively regulated by P1. GOC:dos Process(P2) is directly postively regulated by process(P1) iff: P1 positively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding positively regulates the kinase activity (P2) of protein B then P2 is directly postively regulated by P1. dos 2017-09-17T13:52:47Z directly positively regulated by Process(P2) is directly postively regulated by process(P1) iff: P1 positively regulates P2 via direct physical interaction between an agent executing P1 (or some part of P1) and an agent executing P2 (or some part of P2). For example, if protein A has protein binding activity(P1) that targets protein B and this binding positively regulates the kinase activity (P2) of protein B then P2 is directly postively regulated by P1. GOC:dos A 'has effector activity' B if A and B are GO molecular functions (GO_0003674), A 'has component activity' B and B is the effector (output function) of B. Each compound function has only one effector activity. dos 2017-09-22T14:14:36Z This relation is designed for constructing compound molecular functions, typically in combination with one or more regulatory component activity relations. has effector activity A 'has effector activity' B if A and B are GO molecular functions (GO_0003674), A 'has component activity' B and B is the effector (output function) of B. Each compound function has only one effector activity. GOC:dos David Osumi-Sutherland X ends_after Y iff: end(Y) before_or_simultaneous_with end(X) ends after David Osumi-Sutherland starts_at_end_of X immediately_preceded_by Y iff: end(X) simultaneous_with start(Y) immediately preceded by David Osumi-Sutherland ends_at_start_of meets X immediately_precedes_Y iff: end(X) simultaneous_with start(Y) immediately precedes x overlaps y if and only if there exists some z such that x has part z and z part of y http://purl.obolibrary.org/obo/BFO_0000051 some (http://purl.obolibrary.org/obo/BFO_0000050 some ?Y) overlaps true w 'has component' p if w 'has part' p and w is such that it can be directly disassembled into into n parts p, p2, p3, ..., pn, where these parts are of similar type. The definition of 'has component' is still under discussion. The challenge is in providing a definition that does not imply transitivity. For use in recording has_part with a cardinality constraint, because OWL does not permit cardinality constraints to be used in combination with transitive object properties. In situations where you would want to say something like 'has part exactly 5 digit, you would instead use has_component exactly 5 digit. has component x develops from y if and only if either (a) x directly develops from y or (b) there exists some z such that x directly develops from z and z develops from y Chris Mungall David Osumi-Sutherland Melissa Haendel Terry Meehan RO:0002202 worm_anatomy develops_from develops_from This is the transitive form of the develops from relation develops from develops_from inverse of develops from Chris Mungall David Osumi-Sutherland Terry Meehan develops into p regulates q iff p is causally upstream of q, the execution of p is not constant and varies according to specific conditions, and p influences the rate or magnitude of execution of q due to an effect either on some enabler of q or some enabler of a part of q. Chris Mungall David Hill Tanya Berardini GO Regulation precludes parthood; the regulatory process may not be within the regulated process. regulates (processual) false regulates p negatively regulates q iff p regulates q, and p decreases the rate or magnitude of execution of q. Chris Mungall negatively regulates (process to process) negatively regulates p positively regulates q iff p regulates q, and p increases the rate or magnitude of execution of q. Chris Mungall positively regulates (process to process) positively regulates mechanosensory neuron capable of detection of mechanical stimulus involved in sensory perception (GO:0050974) osteoclast SubClassOf 'capable of' some 'bone resorption' A relation between a material entity (such as a cell) and a process, in which the material entity has the ability to carry out the process. Chris Mungall has function realized in For compatibility with BFO, this relation has a shortcut definition in which the expression "capable of some P" expands to "bearer_of (some realized_by only P)". capable of c stands in this relationship to p if and only if there exists some p' such that c is capable_of p', and p' is part_of p. Chris Mungall has function in capable of part of true Chris Mungall Do not use this relation directly. It is ended as a grouping for relations between occurrents involving the relative timing of their starts and ends. https://docs.google.com/document/d/1kBv1ep_9g3sTR-SD3jqzFqhuwo9TPNF-l-9fUDbO6rM/edit?pli=1 A relation that holds between two occurrents. This is a grouping relation that collects together all the Allen relations. temporally related to p has input c iff: p is a process, c is a material entity, c is a participant in p, c is present at the start of p, and the state of c is modified during p. Chris Mungall consumes has input Mammalian thymus has developmental contribution from some pharyngeal pouch 3; Mammalian thymus has developmental contribution from some pharyngeal pouch 4 [Kardong] x has developmental contribution from y iff x has some part z such that z develops from y Chris Mungall has developmental contribution from inverse of has developmental contribution from Chris Mungall developmentally contributes to Candidate definition: x developmentally related to y if and only if there exists some developmental process (GO:0032502) p such that x and y both participates in p, and x is the output of p and y is the input of p false Chris Mungall In general you should not use this relation to make assertions - use one of the more specific relations below this one This relation groups together various other developmental relations. It is fairly generic, encompassing induction, developmental contribution and direct and transitive develops from developmentally preceded by A faulty traffic light (material entity) whose malfunctioning (a process) is causally upstream of a traffic collision (a process): the traffic light acts upstream of the collision. c acts upstream of p if and only if c enables some f that is involved in p' and p' occurs chronologically before p, is not part of p, and affects the execution of p. c is a material entity and f, p, p' are processes. acts upstream of A gene product that has some activity, where that activity may be a part of a pathway or upstream of the pathway. c acts upstream of or within p if c is enables f, and f is causally upstream of or within p. c is a material entity and p is an process. affects acts upstream of or within Inverse of developmentally preceded by Chris Mungall developmentally succeeded by p is causally upstream of, positive effect q iff p is casually upstream of q, and the execution of p is required for the execution of q. cjm holds between x and y if and only if x is causally upstream of y and the progression of x increases the frequency, rate or extent of y causally upstream of, positive effect p is causally upstream of, negative effect q iff p is casually upstream of q, and the execution of p decreases the execution of q. cjm causally upstream of, negative effect q characteristic of part of w if and only if there exists some p such that q inheres in p and p part of w. Because part_of is transitive, inheres in is a sub-relation of characteristic of part of Chris Mungall inheres in part of characteristic of part of true A mereological relationship or a topological relationship Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving parthood or connectivity relationships mereotopologically related to A relationship that holds between entities participating in some developmental process (GO:0032502) Chris Mungall Do not use this relation directly. It is ended as a grouping for a diverse set of relations, all involving organismal development developmentally related to a particular instances of akt-2 enables some instance of protein kinase activity c enables p iff c is capable of p and c acts to execute p. Chris Mungall catalyzes executes has is catalyzing is executing This relation differs from the parent relation 'capable of' in that the parent is weaker and only expresses a capability that may not be actually realized, whereas this relation is always realized. enables A grouping relationship for any relationship directly involving a function, or that holds because of a function of one of the related entities. Chris Mungall This is a grouping relation that collects relations used for the purpose of connecting structure and function functionally related to this relation holds between c and p when c is part of some c', and c' is capable of p. Chris Mungall false part of structure that is capable of true c involved_in p if and only if c enables some process p', and p' is part of p Chris Mungall actively involved in enables part of involved in inverse of enables Chris Mungall enabled by inverse of regulates Chris Mungall regulated by (processual) regulated by inverse of negatively regulates Chris Mungall negatively regulated by inverse of positively regulates Chris Mungall positively regulated by inverse of has input Chris Mungall input of x has developmental potential involving y iff x is capable of a developmental process with output y. y may be the successor of x, or may be a different structure in the vicinity (as for example in the case of developmental induction). Chris Mungall has developmental potential involving x has potential to developmentrally contribute to y iff x developmentally contributes to y or x is capable of developmentally contributing to y Chris Mungall has potential to developmentally contribute to x has the potential to develop into y iff x develops into y or if x is capable of developing into y Chris Mungall has potential to develop into x has potential to directly develop into y iff x directly develops into y or x is capable of directly developing into y Chris Mungall has potential to directly develop into inverse of upstream of Chris Mungall causally downstream of Chris Mungall immediately causally downstream of p indirectly positively regulates q iff p is indirectly causally upstream of q and p positively regulates q. Chris Mungall indirectly activates indirectly positively regulates p indirectly negatively regulates q iff p is indirectly causally upstream of q and p negatively regulates q. Chris Mungall indirectly inhibits indirectly negatively regulates relation that links two events, processes, states, or objects such that one event, process, state, or object (a cause) contributes to the production of another event, process, state, or object (an effect) where the cause is partly or wholly responsible for the effect, and the effect is partly or wholly dependent on the cause. This branch of the ontology deals with causal relations between entities. It is divided into two branches: causal relations between occurrents/processes, and causal relations between material entities. We take an 'activity flow-centric approach', with the former as primary, and define causal relations between material entities in terms of causal relations between occurrents. To define causal relations in an activity-flow type network, we make use of 3 primitives: * Temporal: how do the intervals of the two occurrents relate? * Is the causal relation regulatory? * Is the influence positive or negative? The first of these can be formalized in terms of the Allen Interval Algebra. Informally, the 3 bins we care about are 'direct', 'indirect' or overlapping. Note that all causal relations should be classified under a RO temporal relation (see the branch under 'temporally related to'). Note that all causal relations are temporal, but not all temporal relations are causal. Two occurrents can be related in time without being causally connected. We take causal influence to be primitive, elucidated as being such that has the upstream changed, some qualities of the donwstream would necessarily be modified. For the second, we consider a relationship to be regulatory if the system in which the activities occur is capable of altering the relationship to achieve some objective. This could include changing the rate of production of a molecule. For the third, we consider the effect of the upstream process on the output(s) of the downstream process. If the level of output is increased, or the rate of production of the output is increased, then the direction is increased. Direction can be positive, negative or neutral or capable of either direction. Two positives in succession yield a positive, two negatives in succession yield a positive, otherwise the default assumption is that the net effect is canceled and the influence is neutral. Each of these 3 primitives can be composed to yield a cross-product of different relation types. Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causally related to relation that links two events, processes, states, or objects such that one event, process, state, or object (a cause) contributes to the production of another event, process, state, or object (an effect) where the cause is partly or wholly responsible for the effect, and the effect is partly or wholly dependent on the cause. https://en.wikipedia.org/wiki/Causality p is causally upstream of q iff p is causally related to q, the end of p precedes the end of q, and p is not an occurrent part of q. Chris Mungall causally upstream of p is immediately causally upstream of q iff p is causally upstream of q, and the end of p is coincident with the beginning of q. Chris Mungall immediately causally upstream of p is 'causally upstream or within' q iff p is causally related to q, and the end of p precedes, or is coincident with, the end of q. We would like to make this disjoint with 'preceded by', but this is prohibited in OWL2 Chris Mungall influences (processual) affects causally upstream of or within inverse of causally upstream of or within Chris Mungall causally downstream of or within c involved in regulation of p if c is involved in some p' and p' regulates some p Chris Mungall involved in regulation of c involved in regulation of p if c is involved in some p' and p' positively regulates some p Chris Mungall involved in positive regulation of c involved in regulation of p if c is involved in some p' and p' negatively regulates some p Chris Mungall involved in negative regulation of c involved in or regulates p if and only if either (i) c is involved in p or (ii) c is involved in regulation of p OWL does not allow defining object properties via a Union Chris Mungall involved in or reguates involved in or involved in regulation of A relationship that holds between two entities in which the processes executed by the two entities are causally connected. Considering relabeling as 'pairwise interacts with' This relation and all sub-relations can be applied to either (1) pairs of entities that are interacting at any moment of time (2) populations or species of entity whose members have the disposition to interact (3) classes whose members have the disposition to interact. Chris Mungall Note that this relationship type, and sub-relationship types may be redundant with process terms from other ontologies. For example, the symbiotic relationship hierarchy parallels GO. The relations are provided as a convenient shortcut. Consider using the more expressive processual form to capture your data. In the future, these relations will be linked to their cognate processes through rules. in pairwise interaction with interacts with http://purl.obolibrary.org/obo/ro/docs/interaction-relations/ http://purl.obolibrary.org/obo/MI_0914 An interaction relationship in which the two partners are molecular entities that directly physically interact with each other for example via a stable binding interaction or a brief interaction during which one modifies the other. Chris Mungall binds molecularly binds with molecularly interacts with http://purl.obolibrary.org/obo/MI_0915 Axiomatization to GO to be added later Chris Mungall An interaction relation between x and y in which x catalyzes a reaction in which a phosphate group is added to y. phosphorylates The entity A, immediately upstream of the entity B, has an activity that regulates an activity performed by B. For example, A and B may be gene products and binding of B by A regulates the kinase activity of B. A and B can be physically interacting but not necessarily. Immediately upstream means there are no intermediate entity between A and B. Chris Mungall Vasundra Touré molecularly controls directly regulates activity of The entity A, immediately upstream of the entity B, has an activity that negatively regulates an activity performed by B. For example, A and B may be gene products and binding of B by A negatively regulates the kinase activity of B. Chris Mungall Vasundra Touré directly inhibits molecularly decreases activity of directly negatively regulates activity of The entity A, immediately upstream of the entity B, has an activity that positively regulates an activity performed by B. For example, A and B may be gene products and binding of B by A positively regulates the kinase activity of B. Chris Mungall Vasundra Touré directly activates molecularly increases activity of directly positively regulates activity of Chris Mungall This property or its subproperties is not to be used directly. These properties exist as helper properties that are used to support OWL reasoning. helper property (not for use in curation) ABal nucleus child nucleus of ABa nucleus (in C elegans) c is a child nucleus of d if and only if c and d are both nuclei and parts of cells c' and d', where c' is derived from d' by mitosis and the genetic material in c is a copy of the generic material in d Chris Mungall This relation is primarily used in the worm anatomy ontology for representing lineage at the level of nuclei. However, it is applicable to any organismal cell lineage. RO:0002476 worm_anatomy DESCENDENTOF DESCENDENTOF child nucleus of A child nucleus relationship in which the cells are part of a hermaphroditic organism Chris Mungall RO:0002477 worm_anatomy DESCINHERM DESCINHERM child nucleus of in hermaphrodite A child nucleus relationship in which the cells are part of a male organism Chris Mungall RO:0002478 worm_anatomy DESCINMALE DESCINMALE child nucleus of in male Chris Mungall is kinase activity A relationship between a material entity and a process where the material entity has some causal role that influences the process causal agent in process p is causally related to q if and only if p or any part of p and q or any part of q are linked by a chain of events where each event pair is one where the execution of p influences the execution of q. p may be upstream, downstream, part of, or a container of q. Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causal relation between processes Chris Mungall depends on The intent is that the process branch of the causal property hierarchy is primary (causal relations hold between occurrents/processes), and that the material branch is defined in terms of the process branch Chris Mungall Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. causal relation between entities Chris Mungall causally influenced by (entity-centric) causally influenced by Chris Mungall interaction relation helper property http://purl.obolibrary.org/obo/ro/docs/interaction-relations/ Chris Mungall molecular interaction relation helper property The entity or characteristic A is causally upstream of the entity or characteristic B, A having an effect on B. An entity corresponds to any biological type of entity as long as a mass is measurable. A characteristic corresponds to a particular specificity of an entity (e.g., phenotype, shape, size). Chris Mungall Vasundra Touré causally influences (entity-centric) causally influences p directly regulates q iff p is immediately causally upstream of q and p regulates q. Chris Mungall directly regulates (processual) directly regulates gland SubClassOf 'has part structure that is capable of' some 'secretion by cell' s 'has part structure that is capable of' p if and only if there exists some part x such that s 'has part' x and x 'capable of' p Chris Mungall has part structure that is capable of A relationship that holds between a material entity and a process in which causality is involved, with either the material entity or some part of the material entity exerting some influence over the process, or the process influencing some aspect of the material entity. Do not use this relation directly. It is intended as a grouping for a diverse set of relations, all involving cause and effect. Chris Mungall causal relation between material entity and a process pyrethroid -> growth Holds between c and p if and only if c is capable of some activity a, and a regulates p. capable of regulating Holds between c and p if and only if c is capable of some activity a, and a negatively regulates p. capable of negatively regulating renin -> arteriolar smooth muscle contraction Holds between c and p if and only if c is capable of some activity a, and a positively regulates p. capable of positively regulating Inverse of 'causal agent in process' process has causal agent p directly positively regulates q iff p is immediately causally upstream of q, and p positively regulates q. directly positively regulates (process to process) directly positively regulates p directly negatively regulates q iff p is immediately causally upstream of q, and p negatively regulates q. directly negatively regulates (process to process) directly negatively regulates Holds between an entity and an process P where the entity enables some larger compound process, and that larger process has-part P. cjm 2018-01-25T23:20:13Z enables subfunction cjm 2018-01-26T23:49:30Z acts upstream of or within, positive effect cjm 2018-01-26T23:49:51Z acts upstream of or within, negative effect c 'acts upstream of, positive effect' p if c is enables f, and f is causally upstream of p, and the direction of f is positive cjm 2018-01-26T23:53:14Z acts upstream of, positive effect c 'acts upstream of, negative effect' p if c is enables f, and f is causally upstream of p, and the direction of f is negative cjm 2018-01-26T23:53:22Z acts upstream of, negative effect cjm 2018-03-13T23:55:05Z causally upstream of or within, negative effect cjm 2018-03-13T23:55:19Z causally upstream of or within, positive effect The entity A has an activity that regulates an activity of the entity B. For example, A and B are gene products where the catalytic activity of A regulates the kinase activity of B. Vasundra Touré regulates activity of p is indirectly causally upstream of q iff p is causally upstream of q and there exists some process r such that p is causally upstream of r and r is causally upstream of q. pg 2022-09-26T06:07:17Z indirectly causally upstream of p indirectly regulates q iff p is indirectly causally upstream of q and p regulates q. pg 2022-09-26T06:08:01Z indirectly regulates A diagnostic testing device utilizes a specimen. X device utilizes material Y means X and Y are material entities, and X is capable of some process P that has input Y. https://orcid.org/0000-0001-9625-1899 https://orcid.org/0000-0003-2620-0345 A diagnostic testing device utilizes a specimen means that the diagnostic testing device is capable of an assay, and this assay a specimen as its input. See github ticket https://github.com/oborel/obo-relations/issues/497 2021-11-08T12:00:00Z utilizes device utilizes material A relationship that holds between a process and a characteristic in which process (P) regulates characteristic (C) iff: P results in the existence of C OR affects the intensity or magnitude of C. regulates characteristic A relationship that holds between a process and a characteristic in which process (P) positively regulates characteristic (C) iff: P results in an increase in the intensity or magnitude of C. positively regulates characteristic A relationship that holds between a process and a characteristic in which process (P) negatively regulates characteristic (C) iff: P results in a decrease in the intensity or magnitude of C. negatively regulates characteristic T1 xunion_of T2 means that T1 is the union of T2 and one (or more) other term, exclusively. Thus, given T1 xunion_of T2 and T1 xunion_T3, one can infer that any instance of T1 must be an instance of T2 or T3 but not T2 and T3. worm_anatomy xunion_of exclusive_union_of T1 xunion_of T2 means that T1 is the union of T2 and one (or more) other term, exclusively. Thus, given T1 xunion_of T2 and T1 xunion_T3, one can infer that any instance of T1 must be an instance of T2 or T3 but not T2 and T3. WB:rynl An entity that exists in full at any time in which it exists at all, persists through time while maintaining its identity and has no temporal parts. continuant An entity that has temporal parts and that happens, unfolds or develops through time. occurrent A continuant that is a bearer of quality and realizable entity entities, in which other entities inhere and which itself cannot inhere in anything. independent continuant An occurrent that has temporal proper parts and for some time t, p s-depends_on some material entity at t. process disposition A specifically dependent continuant that inheres in continuant entities and are not exhibited in full at every time in which it inheres in an entity or group of entities. The exhibition or actualization of a realizable entity is a particular manifestation, functioning or process that occurs under certain circumstances. realizable entity quality A continuant that inheres in or is borne by other entities. Every instance of A requires some specific instance of B which must always be the same. specifically dependent continuant A realizable entity the manifestation of which brings about some result or end that is not essential to a continuant in virtue of the kind of thing that it is but that can be served or participated in by that kind of continuant in some kinds of natural, social or institutional contexts. role function An independent continuant that is spatially extended whose identity is independent of that of other entities and can be maintained through time. material entity anatomical entity biological entity molecular process molecular_function catalytic activity nucleus biological_process true kinase activity transferase activity transferase activity, transferring phosphorus-containing groups quality (PATO) quality An anatomical entity of the nematode Caenorhabditis elegans. worm_anatomy WBbt:0000100 C. elegans anatomical entity An anatomical entity of the nematode Caenorhabditis elegans. WB:WBPerson2987 pedigree of nuclei, lineal descent from P0, the founder. WB:RYNL worm_anatomy WBbt:0000101 Lineage pedigree of nuclei, lineal descent from P0, the founder. WB:RYNL nucleus of pedigree P0 worm_anatomy WBbt:0000102 P0 nucleus nucleus of pedigree P0 WB:rynl nucleus of embryonic cell AB. worm_anatomy WBbt:0001001 AB nucleus nucleus of embryonic cell AB. WB:rynl nucleus of pedigree ABa worm_anatomy WBbt:0001002 ABa nucleus nucleus of pedigree ABa WB:rynl nucleus of pedigree ABal worm_anatomy WBbt:0001003 ABal nucleus nucleus of pedigree ABal WB:rynl nucleus of pedigree ABala worm_anatomy WBbt:0001004 ABala nucleus nucleus of pedigree ABala WB:rynl nucleus of pedigree ABalaa worm_anatomy WBbt:0001005 ABalaa nucleus nucleus of pedigree ABalaa WB:rynl nucleus of pedigree ABalaaa worm_anatomy WBbt:0001006 ABalaaa nucleus nucleus of pedigree ABalaaa WB:rynl nucleus of pedigree ABalaaaa worm_anatomy WBbt:0001007 ABalaaaa nucleus nucleus of pedigree ABalaaaa WB:rynl nucleus of pedigree ABalaaaal worm_anatomy WBbt:0001008 ABalaaaal nucleus nucleus of pedigree ABalaaaal WB:rynl nucleus of pedigree ABalaaaala worm_anatomy WBbt:0001009 ABalaaaala nucleus nucleus of pedigree ABalaaaala WB:rynl nucleus of pedigree ABalaaaalar worm_anatomy WBbt:0001010 ABalaaaalar nucleus nucleus of pedigree ABalaaaalar WB:rynl nucleus of pedigree ABalaaaalp worm_anatomy WBbt:0001011 ABalaaaalp nucleus nucleus of pedigree ABalaaaalp WB:rynl nucleus of pedigree ABalaaaalpa worm_anatomy WBbt:0001012 ABalaaaalpa nucleus nucleus of pedigree ABalaaaalpa WB:rynl nucleus of pedigree ABalaaaar worm_anatomy WBbt:0001013 ABalaaaar nucleus nucleus of pedigree ABalaaaar WB:rynl nucleus of pedigree ABalaaaarl worm_anatomy WBbt:0001014 ABalaaaarl nucleus nucleus of pedigree ABalaaaarl WB:rynl nucleus of pedigree ABalaaaarla worm_anatomy WBbt:0001015 ABalaaaarla nucleus nucleus of pedigree ABalaaaarla WB:rynl nucleus of pedigree ABalaaaarr worm_anatomy WBbt:0001016 ABalaaaarr nucleus nucleus of pedigree ABalaaaarr WB:rynl nucleus of pedigree ABalaaaarra worm_anatomy WBbt:0001017 ABalaaaarra nucleus nucleus of pedigree ABalaaaarra WB:rynl nucleus of pedigree ABalaaap worm_anatomy WBbt:0001018 ABalaaap nucleus nucleus of pedigree ABalaaap WB:rynl nucleus of pedigree ABalaaapa worm_anatomy WBbt:0001019 ABalaaapa nucleus nucleus of pedigree ABalaaapa WB:rynl nucleus of pedigree ABalaaapal worm_anatomy WBbt:0001020 ABalaaapal nucleus nucleus of pedigree ABalaaapal WB:rynl nucleus of pedigree ABalaaapar worm_anatomy WBbt:0001021 ABalaaapar nucleus nucleus of pedigree ABalaaapar WB:rynl nucleus of pedigree ABalaaaparl worm_anatomy WBbt:0001022 ABalaaaparl nucleus nucleus of pedigree ABalaaaparl WB:rynl nucleus of pedigree ABalaaapp worm_anatomy WBbt:0001023 ABalaaapp nucleus nucleus of pedigree ABalaaapp WB:rynl nucleus of pedigree ABalaaappl worm_anatomy WBbt:0001024 ABalaaappl nucleus nucleus of pedigree ABalaaappl WB:rynl nucleus of pedigree ABalaaapplr worm_anatomy WBbt:0001025 ABalaaapplr nucleus nucleus of pedigree ABalaaapplr WB:rynl nucleus of pedigree ABalaaappr worm_anatomy WBbt:0001026 ABalaaappr nucleus nucleus of pedigree ABalaaappr WB:rynl nucleus of pedigree ABalaap worm_anatomy WBbt:0001027 ABalaap nucleus nucleus of pedigree ABalaap WB:rynl nucleus of pedigree ABalaapa worm_anatomy WBbt:0001028 ABalaapa nucleus nucleus of pedigree ABalaapa WB:rynl nucleus of pedigree ABalaapaa worm_anatomy WBbt:0001029 ABalaapaa nucleus nucleus of pedigree ABalaapaa WB:rynl nucleus of pedigree ABalaapaaa worm_anatomy WBbt:0001030 ABalaapaaa nucleus nucleus of pedigree ABalaapaaa WB:rynl nucleus of pedigree ABalaapaaal worm_anatomy WBbt:0001031 ABalaapaaal nucleus nucleus of pedigree ABalaapaaal WB:rynl nucleus of pedigree ABalaapaap worm_anatomy WBbt:0001032 ABalaapaap nucleus nucleus of pedigree ABalaapaap WB:rynl nucleus of pedigree ABalaapaapa worm_anatomy WBbt:0001033 ABalaapaapa nucleus nucleus of pedigree ABalaapaapa WB:rynl nucleus of pedigree ABalaapap worm_anatomy WBbt:0001034 ABalaapap nucleus nucleus of pedigree ABalaapap WB:rynl nucleus of pedigree ABalaapapa worm_anatomy WBbt:0001035 ABalaapapa nucleus nucleus of pedigree ABalaapapa WB:rynl nucleus of pedigree ABalaapapp worm_anatomy WBbt:0001036 ABalaapapp nucleus nucleus of pedigree ABalaapapp WB:rynl nucleus of pedigree ABalaapappa worm_anatomy WBbt:0001037 ABalaapappa nucleus nucleus of pedigree ABalaapappa WB:rynl nucleus of pedigree ABalaapp worm_anatomy WBbt:0001038 ABalaapp nucleus nucleus of pedigree ABalaapp WB:rynl nucleus of pedigree ABalaappa worm_anatomy WBbt:0001039 ABalaappa nucleus nucleus of pedigree ABalaappa WB:rynl nucleus of pedigree ABalaappaa worm_anatomy WBbt:0001040 ABalaappaa nucleus nucleus of pedigree ABalaappaa WB:rynl nucleus of pedigree ABalaappap worm_anatomy WBbt:0001041 ABalaappap nucleus nucleus of pedigree ABalaappap WB:rynl nucleus of pedigree ABalaappp worm_anatomy WBbt:0001042 ABalaappp nucleus nucleus of pedigree ABalaappp WB:rynl nucleus of pedigree ABalaapppa worm_anatomy WBbt:0001043 ABalaapppa nucleus nucleus of pedigree ABalaapppa WB:rynl nucleus of pedigree ABalaapppap worm_anatomy WBbt:0001044 ABalaapppap nucleus nucleus of pedigree ABalaapppap WB:rynl nucleus of pedigree ABalaapppp worm_anatomy WBbt:0001045 ABalaapppp nucleus nucleus of pedigree ABalaapppp WB:rynl nucleus of pedigree ABalaappppa worm_anatomy WBbt:0001046 ABalaappppa nucleus nucleus of pedigree ABalaappppa WB:rynl nucleus of pedigree ABalaappppap worm_anatomy WBbt:0001047 ABalaappppap nucleus nucleus of pedigree ABalaappppap WB:rynl nucleus of pedigree ABalap worm_anatomy WBbt:0001048 ABalap nucleus nucleus of pedigree ABalap WB:rynl nucleus of pedigree ABalapa worm_anatomy WBbt:0001049 ABalapa nucleus nucleus of pedigree ABalapa WB:rynl nucleus of pedigree ABalapaa worm_anatomy WBbt:0001050 ABalapaa nucleus nucleus of pedigree ABalapaa WB:rynl nucleus of pedigree ABalapaaa worm_anatomy WBbt:0001051 ABalapaaa nucleus nucleus of pedigree ABalapaaa WB:rynl nucleus of pedigree ABalapaaaa worm_anatomy WBbt:0001052 ABalapaaaa nucleus nucleus of pedigree ABalapaaaa WB:rynl nucleus of pedigree ABalapaaaap worm_anatomy WBbt:0001053 ABalapaaaap nucleus nucleus of pedigree ABalapaaaap WB:rynl nucleus of pedigree ABalapaaap worm_anatomy WBbt:0001054 ABalapaaap nucleus nucleus of pedigree ABalapaaap WB:rynl nucleus of pedigree ABalapaap worm_anatomy WBbt:0001055 ABalapaap nucleus nucleus of pedigree ABalapaap WB:rynl nucleus of pedigree ABalapaapa worm_anatomy WBbt:0001056 ABalapaapa nucleus nucleus of pedigree ABalapaapa WB:rynl nucleus of pedigree ABalapaapap worm_anatomy WBbt:0001057 ABalapaapap nucleus nucleus of pedigree ABalapaapap WB:rynl nucleus of pedigree ABalapaapp worm_anatomy WBbt:0001058 ABalapaapp nucleus nucleus of pedigree ABalapaapp WB:rynl nucleus of pedigree ABalapaappa worm_anatomy WBbt:0001059 ABalapaappa nucleus nucleus of pedigree ABalapaappa WB:rynl nucleus of pedigree ABalapaappap worm_anatomy WBbt:0001060 ABalapaappap nucleus nucleus of pedigree ABalapaappap WB:rynl nucleus of pedigree ABalapap worm_anatomy WBbt:0001061 ABalapap nucleus nucleus of pedigree ABalapap WB:rynl nucleus of pedigree ABalapapa worm_anatomy WBbt:0001062 ABalapapa nucleus nucleus of pedigree ABalapapa WB:rynl nucleus of pedigree ABalapapaa worm_anatomy WBbt:0001063 ABalapapaa nucleus nucleus of pedigree ABalapapaa WB:rynl nucleus of pedigree ABalapapap worm_anatomy WBbt:0001064 ABalapapap nucleus nucleus of pedigree ABalapapap WB:rynl nucleus of pedigree ABalapapapa worm_anatomy WBbt:0001065 ABalapapapa nucleus nucleus of pedigree ABalapapapa WB:rynl nucleus of pedigree ABalapapapap worm_anatomy WBbt:0001066 ABalapapapap nucleus nucleus of pedigree ABalapapapap WB:rynl nucleus of pedigree ABalapapp worm_anatomy WBbt:0001067 ABalapapp nucleus nucleus of pedigree ABalapapp WB:rynl nucleus of pedigree ABalapappa worm_anatomy WBbt:0001068 ABalapappa nucleus nucleus of pedigree ABalapappa WB:rynl nucleus of pedigree ABalapappaa worm_anatomy WBbt:0001069 ABalapappaa nucleus nucleus of pedigree ABalapappaa WB:rynl nucleus of pedigree ABalapappaap worm_anatomy WBbt:0001070 ABalapappaap nucleus nucleus of pedigree ABalapappaap WB:rynl nucleus of pedigree ABalapappp worm_anatomy WBbt:0001071 ABalapappp nucleus nucleus of pedigree ABalapappp WB:rynl nucleus of pedigree ABalapp worm_anatomy WBbt:0001072 ABalapp nucleus nucleus of pedigree ABalapp WB:rynl nucleus of pedigree ABalappa worm_anatomy WBbt:0001073 ABalappa nucleus nucleus of pedigree ABalappa WB:rynl nucleus of pedigree ABalappaa worm_anatomy WBbt:0001074 ABalappaa nucleus nucleus of pedigree ABalappaa WB:rynl nucleus of pedigree ABalappaaa worm_anatomy WBbt:0001075 ABalappaaa nucleus nucleus of pedigree ABalappaaa WB:rynl nucleus of pedigree ABalappaap worm_anatomy WBbt:0001076 ABalappaap nucleus nucleus of pedigree ABalappaap WB:rynl nucleus of pedigree ABalappaapp worm_anatomy WBbt:0001077 ABalappaapp nucleus nucleus of pedigree ABalappaapp WB:rynl nucleus of pedigree ABalappap worm_anatomy WBbt:0001078 ABalappap nucleus nucleus of pedigree ABalappap WB:rynl nucleus of pedigree ABalappapa worm_anatomy WBbt:0001079 ABalappapa nucleus nucleus of pedigree ABalappapa WB:rynl nucleus of pedigree ABalappapap worm_anatomy WBbt:0001080 ABalappapap nucleus nucleus of pedigree ABalappapap WB:rynl nucleus of pedigree ABalappapp worm_anatomy WBbt:0001081 ABalappapp nucleus nucleus of pedigree ABalappapp WB:rynl nucleus of pedigree ABalappp worm_anatomy WBbt:0001082 ABalappp nucleus nucleus of pedigree ABalappp WB:rynl nucleus of pedigree ABalapppa worm_anatomy WBbt:0001083 ABalapppa nucleus nucleus of pedigree ABalapppa WB:rynl nucleus of pedigree ABalapppaa worm_anatomy WBbt:0001084 ABalapppaa nucleus nucleus of pedigree ABalapppaa WB:rynl nucleus of pedigree ABalapppap worm_anatomy WBbt:0001085 ABalapppap nucleus nucleus of pedigree ABalapppap WB:rynl nucleus of pedigree ABalapppapa worm_anatomy WBbt:0001086 ABalapppapa nucleus nucleus of pedigree ABalapppapa WB:rynl nucleus of pedigree ABalapppapap worm_anatomy WBbt:0001087 ABalapppapap nucleus nucleus of pedigree ABalapppapap WB:rynl nucleus of pedigree ABalapppp worm_anatomy WBbt:0001088 ABalapppp nucleus nucleus of pedigree ABalapppp WB:rynl nucleus of pedigree ABalappppa worm_anatomy WBbt:0001089 ABalappppa nucleus nucleus of pedigree ABalappppa WB:rynl nucleus of pedigree ABalappppaa worm_anatomy WBbt:0001090 ABalappppaa nucleus nucleus of pedigree ABalappppaa WB:rynl nucleus of pedigree ABalappppaap worm_anatomy WBbt:0001091 ABalappppaap nucleus nucleus of pedigree ABalappppaap WB:rynl nucleus of pedigree ABalappppp worm_anatomy WBbt:0001092 ABalappppp nucleus nucleus of pedigree ABalappppp WB:rynl nucleus of pedigree ABalp worm_anatomy WBbt:0001093 ABalp nucleus nucleus of pedigree ABalp WB:rynl nucleus of pedigree ABalpa worm_anatomy WBbt:0001094 ABalpa nucleus nucleus of pedigree ABalpa WB:rynl nucleus of pedigree ABalpaa worm_anatomy WBbt:0001095 ABalpaa nucleus nucleus of pedigree ABalpaa WB:rynl nucleus of pedigree ABalpaaa worm_anatomy WBbt:0001096 ABalpaaa nucleus nucleus of pedigree ABalpaaa WB:rynl nucleus of pedigree ABalpaaaa worm_anatomy WBbt:0001097 ABalpaaaa nucleus nucleus of pedigree ABalpaaaa WB:rynl nucleus of pedigree ABalpaaaaa worm_anatomy WBbt:0001098 ABalpaaaaa nucleus nucleus of pedigree ABalpaaaaa WB:rynl nucleus of pedigree ABalpaaaap worm_anatomy WBbt:0001099 ABalpaaaap nucleus nucleus of pedigree ABalpaaaap WB:rynl nucleus of pedigree ABalpaaaapp worm_anatomy WBbt:0001100 ABalpaaaapp nucleus nucleus of pedigree ABalpaaaapp WB:rynl nucleus of pedigree ABalpaaap worm_anatomy WBbt:0001101 ABalpaaap nucleus nucleus of pedigree ABalpaaap WB:rynl nucleus of pedigree ABalpaaapa worm_anatomy WBbt:0001102 ABalpaaapa nucleus nucleus of pedigree ABalpaaapa WB:rynl nucleus of pedigree ABalpaaapap worm_anatomy WBbt:0001103 ABalpaaapap nucleus nucleus of pedigree ABalpaaapap WB:rynl nucleus of pedigree ABalpaaapp worm_anatomy WBbt:0001104 ABalpaaapp nucleus nucleus of pedigree ABalpaaapp WB:rynl nucleus of pedigree ABalpaaappa worm_anatomy WBbt:0001105 ABalpaaappa nucleus nucleus of pedigree ABalpaaappa WB:rynl nucleus of pedigree ABalpaap worm_anatomy WBbt:0001106 ABalpaap nucleus nucleus of pedigree ABalpaap WB:rynl nucleus of pedigree ABalpaapa worm_anatomy WBbt:0001107 ABalpaapa nucleus nucleus of pedigree ABalpaapa WB:rynl nucleus of pedigree ABalpaapaa worm_anatomy WBbt:0001108 ABalpaapaa nucleus nucleus of pedigree ABalpaapaa WB:rynl nucleus of pedigree ABalpaapap worm_anatomy WBbt:0001109 ABalpaapap nucleus nucleus of pedigree ABalpaapap WB:rynl nucleus of pedigree ABalpaapp worm_anatomy WBbt:0001110 ABalpaapp nucleus nucleus of pedigree ABalpaapp WB:rynl nucleus of pedigree ABalpaappa worm_anatomy WBbt:0001111 ABalpaappa nucleus nucleus of pedigree ABalpaappa WB:rynl nucleus of pedigree ABalpaappp worm_anatomy WBbt:0001112 ABalpaappp nucleus nucleus of pedigree ABalpaappp WB:rynl nucleus of pedigree ABalpap worm_anatomy WBbt:0001113 ABalpap nucleus nucleus of pedigree ABalpap WB:rynl nucleus of pedigree ABalpapa worm_anatomy WBbt:0001114 ABalpapa nucleus nucleus of pedigree ABalpapa WB:rynl nucleus of pedigree ABalpapaa worm_anatomy WBbt:0001115 ABalpapaa nucleus nucleus of pedigree ABalpapaa WB:rynl nucleus of pedigree ABalpapaaa worm_anatomy WBbt:0001116 ABalpapaaa nucleus nucleus of pedigree ABalpapaaa WB:rynl nucleus of pedigree ABalpapaap worm_anatomy WBbt:0001117 ABalpapaap nucleus nucleus of pedigree ABalpapaap WB:rynl nucleus of pedigree ABalpapaapp worm_anatomy WBbt:0001118 ABalpapaapp nucleus nucleus of pedigree ABalpapaapp WB:rynl nucleus of pedigree ABalpapap worm_anatomy WBbt:0001119 ABalpapap nucleus nucleus of pedigree ABalpapap WB:rynl nucleus of pedigree ABalpapapa worm_anatomy WBbt:0001120 ABalpapapa nucleus nucleus of pedigree ABalpapapa WB:rynl nucleus of pedigree ABalpapapap worm_anatomy WBbt:0001121 ABalpapapap nucleus nucleus of pedigree ABalpapapap WB:rynl nucleus of pedigree ABalpapapapa worm_anatomy WBbt:0001122 ABalpapapapa nucleus nucleus of pedigree ABalpapapapa WB:rynl nucleus of pedigree ABalpapapp worm_anatomy WBbt:0001123 ABalpapapp nucleus nucleus of pedigree ABalpapapp WB:rynl nucleus of pedigree ABalpapappa worm_anatomy WBbt:0001124 ABalpapappa nucleus nucleus of pedigree ABalpapappa WB:rynl nucleus of pedigree ABalpapp worm_anatomy WBbt:0001125 ABalpapp nucleus nucleus of pedigree ABalpapp WB:rynl nucleus of pedigree ABalpappa worm_anatomy WBbt:0001126 ABalpappa nucleus nucleus of pedigree ABalpappa WB:rynl nucleus of pedigree ABalpappaa worm_anatomy WBbt:0001127 ABalpappaa nucleus nucleus of pedigree ABalpappaa WB:rynl nucleus of pedigree ABalpappaaa worm_anatomy WBbt:0001128 ABalpappaaa nucleus nucleus of pedigree ABalpappaaa WB:rynl nucleus of pedigree ABalpappaap worm_anatomy WBbt:0001129 ABalpappaap nucleus nucleus of pedigree ABalpappaap WB:rynl nucleus of pedigree ABalpappaapp worm_anatomy WBbt:0001130 ABalpappaapp nucleus nucleus of pedigree ABalpappaapp WB:rynl nucleus of pedigree ABalpappap worm_anatomy WBbt:0001131 ABalpappap nucleus nucleus of pedigree ABalpappap WB:rynl nucleus of pedigree ABalpappp worm_anatomy WBbt:0001132 ABalpappp nucleus nucleus of pedigree ABalpappp WB:rynl nucleus of pedigree ABalpapppa worm_anatomy WBbt:0001133 ABalpapppa nucleus nucleus of pedigree ABalpapppa WB:rynl nucleus of pedigree ABalpapppap worm_anatomy WBbt:0001134 ABalpapppap nucleus nucleus of pedigree ABalpapppap WB:rynl nucleus of pedigree ABalpapppp worm_anatomy WBbt:0001135 ABalpapppp nucleus nucleus of pedigree ABalpapppp WB:rynl nucleus of pedigree ABalpp worm_anatomy WBbt:0001136 ABalpp nucleus nucleus of pedigree ABalpp WB:rynl nucleus of pedigree ABalppa worm_anatomy WBbt:0001137 ABalppa nucleus nucleus of pedigree ABalppa WB:rynl nucleus of pedigree ABalppaa worm_anatomy WBbt:0001138 ABalppaa nucleus nucleus of pedigree ABalppaa WB:rynl nucleus of pedigree ABalppaaa worm_anatomy WBbt:0001139 ABalppaaa nucleus nucleus of pedigree ABalppaaa WB:rynl nucleus of pedigree ABalppaaaa worm_anatomy WBbt:0001140 ABalppaaaa nucleus nucleus of pedigree ABalppaaaa WB:rynl nucleus of pedigree ABalppaaap worm_anatomy WBbt:0001141 ABalppaaap nucleus nucleus of pedigree ABalppaaap WB:rynl nucleus of pedigree ABalppaap worm_anatomy WBbt:0001142 ABalppaap nucleus nucleus of pedigree ABalppaap WB:rynl nucleus of pedigree ABalppaapa worm_anatomy WBbt:0001143 ABalppaapa nucleus nucleus of pedigree ABalppaapa WB:rynl nucleus of pedigree ABalppaapp worm_anatomy WBbt:0001144 ABalppaapp nucleus nucleus of pedigree ABalppaapp WB:rynl nucleus of pedigree ABalppaappa worm_anatomy WBbt:0001145 ABalppaappa nucleus nucleus of pedigree ABalppaappa WB:rynl nucleus of pedigree ABalppap worm_anatomy WBbt:0001146 ABalppap nucleus nucleus of pedigree ABalppap WB:rynl nucleus of pedigree ABalppapa worm_anatomy WBbt:0001147 ABalppapa nucleus nucleus of pedigree ABalppapa WB:rynl nucleus of pedigree ABalppapaa worm_anatomy WBbt:0001148 ABalppapaa nucleus nucleus of pedigree ABalppapaa WB:rynl nucleus of pedigree ABalppapap worm_anatomy WBbt:0001149 ABalppapap nucleus nucleus of pedigree ABalppapap WB:rynl nucleus of pedigree ABalppapp worm_anatomy WBbt:0001150 ABalppapp nucleus nucleus of pedigree ABalppapp WB:rynl nucleus of pedigree ABalppappa worm_anatomy WBbt:0001151 ABalppappa nucleus nucleus of pedigree ABalppappa WB:rynl nucleus of pedigree ABalppappp worm_anatomy WBbt:0001152 ABalppappp nucleus nucleus of pedigree ABalppappp WB:rynl nucleus of pedigree ABalppapppa worm_anatomy WBbt:0001153 ABalppapppa nucleus nucleus of pedigree ABalppapppa WB:rynl nucleus of pedigree ABalppapppap worm_anatomy WBbt:0001154 ABalppapppap nucleus nucleus of pedigree ABalppapppap WB:rynl nucleus of pedigree ABalppp worm_anatomy WBbt:0001155 ABalppp nucleus nucleus of pedigree ABalppp WB:rynl nucleus of pedigree ABalpppa worm_anatomy WBbt:0001156 ABalpppa nucleus nucleus of pedigree ABalpppa WB:rynl nucleus of pedigree ABalpppaa worm_anatomy WBbt:0001157 ABalpppaa nucleus nucleus of pedigree ABalpppaa WB:rynl nucleus of pedigree ABalpppaaa worm_anatomy WBbt:0001158 ABalpppaaa nucleus nucleus of pedigree ABalpppaaa WB:rynl nucleus of pedigree ABalpppaap worm_anatomy WBbt:0001159 ABalpppaap nucleus nucleus of pedigree ABalpppaap WB:rynl nucleus of pedigree ABalpppaapv worm_anatomy WBbt:0001160 ABalpppaapv nucleus nucleus of pedigree ABalpppaapv WB:rynl nucleus of pedigree ABalpppap worm_anatomy WBbt:0001161 ABalpppap nucleus nucleus of pedigree ABalpppap WB:rynl nucleus of pedigree ABalpppapa worm_anatomy WBbt:0001162 ABalpppapa nucleus nucleus of pedigree ABalpppapa WB:rynl nucleus of pedigree ABalpppapp worm_anatomy WBbt:0001163 ABalpppapp nucleus nucleus of pedigree ABalpppapp WB:rynl nucleus of pedigree ABalpppappa worm_anatomy WBbt:0001164 ABalpppappa nucleus nucleus of pedigree ABalpppappa WB:rynl nucleus of pedigree ABalpppappp worm_anatomy WBbt:0001165 ABalpppappp nucleus nucleus of pedigree ABalpppappp WB:rynl nucleus of pedigree ABalpppapppp worm_anatomy WBbt:0001166 ABalpppapppp nucleus nucleus of pedigree ABalpppapppp WB:rynl nucleus of pedigree ABalpppp worm_anatomy WBbt:0001167 ABalpppp nucleus nucleus of pedigree ABalpppp WB:rynl nucleus of pedigree ABalppppa worm_anatomy WBbt:0001168 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