@prefix : <http://togoid.dbcls.jp/ontology#> .
@prefix owl: <http://www.w3.org/2002/07/owl#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix dcterms: <http://purl.org/dc/terms/> .
@prefix skos: <http://www.w3.org/2004/02/skos/core#> .

: a owl:Ontology .

:Category
    a owl:Class ;
    rdfs:label "Category" .

:Dataset
    a owl:Class ;
    rdfs:label "Dataset" .

:relation
    a owl:ObjectProperty ;
    rdfs:comment "Identifiers of the domain dataset have a biological relation with that of the range dataset." .

:display_label
    a owl:DatatypeProperty ;
    rdfs:comment "A label to be displayed on the TogoID web UI." .

:Glycan
    a owl:Class ;
    rdfs:label "Glycan" ;
    rdfs:subClassOf :Category , :Molecule .

:Compound
    a owl:Class ;
    rdfs:label "Compound" ;
    rdfs:subClassOf :Category , :Molecule .

:Phenotype
    a owl:Class ;
    rdfs:label "Phenotype" ;
    rdfs:subClassOf :Category .

:Domain
    a owl:Class ;
    rdfs:label "Domain" ;
    rdfs:subClassOf :Category .

:Function
    a owl:Class ;
    rdfs:label "Function" ;
    rdfs:subClassOf :Category .

:Transcript
    a owl:Class ;
    rdfs:label "Transcript" ;
    rdfs:subClassOf :Category , :Sequence .

:Gene
    a owl:Class ;
    rdfs:label "Gene" ;
    rdfs:subClassOf :Category , :Sequence .

:Interaction
    a owl:Class ;
    rdfs:label "Interaction" ;
    rdfs:subClassOf :Category .

:Reaction
    a owl:Class ;
    rdfs:label "Reaction" ;
    rdfs:subClassOf :Category .

:Literature
    a owl:Class ;
    rdfs:label "Literature" ;
    rdfs:subClassOf :Category .

:Submission
    a owl:Class ;
    rdfs:label "Submission" ;
    rdfs:subClassOf :Category .

:Analysis
    a owl:Class ;
    rdfs:label "Analysis" ;
    rdfs:subClassOf :Category .

:Experiment
    a owl:Class ;
    rdfs:label "Experiment" ;
    rdfs:subClassOf :Category .

:SequenceRun
    a owl:Class ;
    rdfs:label "Sequence run" ;
    rdfs:subClassOf :Category .

:Ortholog
    a owl:Class ;
    rdfs:label "Ortholog" ;
    rdfs:subClassOf :Category .

:Pathway
    a owl:Class ;
    rdfs:label "Pathway" ;
    rdfs:subClassOf :Category .

:Probe
    a owl:Class ;
    rdfs:label "Probe" ;
    rdfs:subClassOf :Category .

:Project
    a owl:Class ;
    rdfs:label "Project" ;
    rdfs:subClassOf :Category .

:Protein
    a owl:Class ;
    rdfs:label "Protein" ;
    rdfs:subClassOf :Category , :Sequence , :Molecule .

:Sample
    a owl:Class ;
    rdfs:label "Sample" ;
    rdfs:subClassOf :Category .

:Structure
    a owl:Class ;
    rdfs:label "Structure" ;
    rdfs:subClassOf :Category .

:Organism
    a owl:Class ;
    rdfs:label "Organism" ;
    rdfs:subClassOf :Category .

:Variant
    a owl:Class ;
    rdfs:label "Variant" ;
    rdfs:subClassOf :Category .

:Synonym
    a owl:Class ;
    rdfs:label "Synonym" ;
    rdfs:subClassOf :Category .

:Sequence
    a owl:Class ;
    rdfs:label "Sequence" ;
    rdfs:subClassOf :Category .

:Molecule
    a owl:Class ;
    rdfs:label "Molecule" ;
    rdfs:subClassOf :Category .

:Genome
    a owl:Class ;
    rdfs:label "Genome" ;
    rdfs:subClassOf :Category .

:CellLine
    a owl:Class ;
    rdfs:label "Cell line" ;
    rdfs:subClassOf :Category .

:Anatomy
    a owl:Class ;
    rdfs:label "Anatomy" ;
    rdfs:subClassOf :Category .

:Lipid
    a owl:Class ;
    rdfs:label "Lipid" ;
    rdfs:subClassOf :Category , :Molecule .

:Drug
    a owl:Class ;
    rdfs:label "Drug" ;
    rdfs:subClassOf :Category .

:Proteome
    a owl:Class ;
    rdfs:label "Proteome" ;
    rdfs:subClassOf :Category .

:Classification
    a owl:Class ;
    rdfs:label "Classification" ;
    rdfs:subClassOf :Category .

:AnnotationRule
    a owl:Class ;
    rdfs:label "Annotation rule" ;
    rdfs:subClassOf :Category .

:Genotype
    a owl:Class ;
    rdfs:label "Genotype" ;
    rdfs:subClassOf :Category .

:AffyProbeset
    a owl:Class ;
    dcterms:identifier "affy_probeset" ;
    rdfs:label "Affymetrix probeset" ;
    rdfs:subClassOf :Probe, :Dataset .

:AssemblyInsdc
    a owl:Class ;
    dcterms:identifier "assembly_insdc" ;
    rdfs:label "Assembly INSDC" ;
    rdfs:subClassOf :Genome, :Dataset .

:AssemblyRefseq
    a owl:Class ;
    dcterms:identifier "assembly_refseq" ;
    rdfs:label "Assembly RefSeq" ;
    rdfs:subClassOf :Genome, :Dataset .

:Atc
    a owl:Class ;
    dcterms:identifier "atc" ;
    rdfs:label "ATC classification" ;
    rdfs:subClassOf :Drug, :Dataset .

:Bioproject
    a owl:Class ;
    dcterms:identifier "bioproject" ;
    rdfs:label "BioProject" ;
    rdfs:subClassOf :Project, :Dataset .

:BioprojectUmbrella
    a owl:Class ;
    dcterms:identifier "bioproject_umbrella" ;
    rdfs:label "BioProject umbrella" ;
    rdfs:subClassOf :Project, :Dataset .

:Biosample
    a owl:Class ;
    dcterms:identifier "biosample" ;
    rdfs:label "BioSample" ;
    rdfs:subClassOf :Sample, :Dataset .

:Ccds
    a owl:Class ;
    dcterms:identifier "ccds" ;
    rdfs:label "Consensus CDS" ;
    rdfs:subClassOf :Gene, :Dataset .

:Cellosaurus
    a owl:Class ;
    dcterms:identifier "cellosaurus" ;
    rdfs:label "Cellosaurus" ;
    rdfs:subClassOf :CellLine, :Dataset .

:Chebi
    a owl:Class ;
    dcterms:identifier "chebi" ;
    rdfs:label "ChEBI compound" ;
    rdfs:subClassOf :Compound, :Dataset .

:ChemblCompound
    a owl:Class ;
    dcterms:identifier "chembl_compound" ;
    rdfs:label "ChEMBL compound" ;
    rdfs:subClassOf :Compound, :Dataset .

:ChemblTarget
    a owl:Class ;
    dcterms:identifier "chembl_target" ;
    rdfs:label "ChEMBL target" ;
    rdfs:subClassOf :Protein, :Dataset .

:Clinvar
    a owl:Class ;
    dcterms:identifier "clinvar" ;
    rdfs:label "ClinVar variant" ;
    rdfs:subClassOf :Variant, :Dataset .

:Cog
    a owl:Class ;
    dcterms:identifier "cog" ;
    rdfs:label "COG" ;
    rdfs:subClassOf :Ortholog, :Dataset .

:Dbsnp
    a owl:Class ;
    dcterms:identifier "dbsnp" ;
    rdfs:label "dbSNP" ;
    rdfs:subClassOf :Variant, :Dataset .

:Doid
    a owl:Class ;
    dcterms:identifier "doid" ;
    rdfs:label "Disease ontology" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Drugbank
    a owl:Class ;
    dcterms:identifier "drugbank" ;
    rdfs:label "DrugBank" ;
    rdfs:subClassOf :Drug, :Dataset .

:Ec
    a owl:Class ;
    dcterms:identifier "ec" ;
    rdfs:label "Enzyme nomenclature" ;
    rdfs:subClassOf :Function, :Dataset .

:EnsemblGene
    a owl:Class ;
    dcterms:identifier "ensembl_gene" ;
    rdfs:label "Ensembl gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:EnsemblProtein
    a owl:Class ;
    dcterms:identifier "ensembl_protein" ;
    rdfs:label "Ensembl protein" ;
    rdfs:subClassOf :Protein, :Dataset .

:EnsemblTranscript
    a owl:Class ;
    dcterms:identifier "ensembl_transcript" ;
    rdfs:label "Ensembl transcript" ;
    rdfs:subClassOf :Transcript, :Dataset .

:FlybaseGene
    a owl:Class ;
    dcterms:identifier "flybase_gene" ;
    rdfs:label "FlyBase gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:Gea
    a owl:Class ;
    dcterms:identifier "gea" ;
    rdfs:label "GEA" ;
    rdfs:subClassOf :Experiment, :Dataset .

:GeoSample
    a owl:Class ;
    dcterms:identifier "geo_sample" ;
    rdfs:label "GEO sample" ;
    rdfs:subClassOf :Sample, :Dataset .

:GeoSeries
    a owl:Class ;
    dcterms:identifier "geo_series" ;
    rdfs:label "GEO series" ;
    rdfs:subClassOf :Project, :Dataset .

:Glytoucan
    a owl:Class ;
    dcterms:identifier "glytoucan" ;
    rdfs:label "GlyTouCan" ;
    rdfs:subClassOf :Glycan, :Dataset .

:Go
    a owl:Class ;
    dcterms:identifier "go" ;
    rdfs:label "Gene ontology" ;
    rdfs:subClassOf :Function, :Dataset .

:Hgnc
    a owl:Class ;
    dcterms:identifier "hgnc" ;
    rdfs:label "HGNC" ;
    rdfs:subClassOf :Gene, :Dataset .

:HgncSymbol
    a owl:Class ;
    dcterms:identifier "hgnc_symbol" ;
    rdfs:label "HGNC gene symbol" ;
    rdfs:subClassOf :Gene , :Synonym, :Dataset .

:Hmdb
    a owl:Class ;
    dcterms:identifier "hmdb" ;
    rdfs:label "HMDB" ;
    rdfs:subClassOf :Compound, :Dataset .

:Homologene
    a owl:Class ;
    dcterms:identifier "homologene" ;
    rdfs:label "HomoloGene" ;
    rdfs:subClassOf :Ortholog, :Dataset .

:HpInheritance
    a owl:Class ;
    dcterms:identifier "hp_inheritance" ;
    rdfs:label "HP Inheritance" ;
    rdfs:subClassOf :Classification, :Dataset .

:HpPhenotype
    a owl:Class ;
    dcterms:identifier "hp_phenotype" ;
    rdfs:label "HP Phenotype" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:InchiKey
    a owl:Class ;
    dcterms:identifier "inchi_key" ;
    rdfs:label "InChIKey" ;
    rdfs:subClassOf :Compound, :Dataset .

:Insdc
    a owl:Class ;
    dcterms:identifier "insdc" ;
    rdfs:label "GenBank/ENA/DDBJ" ;
    rdfs:subClassOf :Gene, :Dataset .

:InsdcMaster
    a owl:Class ;
    dcterms:identifier "insdc_master" ;
    rdfs:label "INSDC master" ;
    rdfs:subClassOf :Project, :Dataset .

:Intact
    a owl:Class ;
    dcterms:identifier "intact" ;
    rdfs:label "IntAct" ;
    rdfs:subClassOf :Interaction, :Dataset .

:Interpro
    a owl:Class ;
    dcterms:identifier "interpro" ;
    rdfs:label "InterPro" ;
    rdfs:subClassOf :Domain, :Dataset .

:IupharLigand
    a owl:Class ;
    dcterms:identifier "iuphar_ligand" ;
    rdfs:label "IUPHAR ligand" ;
    rdfs:subClassOf :Compound, :Dataset .

:JgaDataset
    a owl:Class ;
    dcterms:identifier "jga_dataset" ;
    rdfs:label "JGA Dataset" ;
    rdfs:subClassOf :Project, :Dataset .

:JgaStudy
    a owl:Class ;
    dcterms:identifier "jga_study" ;
    rdfs:label "JGA Study" ;
    rdfs:subClassOf :Project, :Dataset .

:Lipidmaps
    a owl:Class ;
    dcterms:identifier "lipidmaps" ;
    rdfs:label "LIPID MAPS" ;
    rdfs:subClassOf :Lipid, :Dataset .

:Lrg
    a owl:Class ;
    dcterms:identifier "lrg" ;
    rdfs:label "LRG" ;
    rdfs:subClassOf :Gene, :Dataset .

:MbgdGene
    a owl:Class ;
    dcterms:identifier "mbgd_gene" ;
    rdfs:label "MBGD gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:MbgdOrganism
    a owl:Class ;
    dcterms:identifier "mbgd_organism" ;
    rdfs:label "MBGD organism" ;
    rdfs:subClassOf :Organism, :Dataset .

:Meddra
    a owl:Class ;
    dcterms:identifier "meddra" ;
    rdfs:label "MedDRA" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Medgen
    a owl:Class ;
    dcterms:identifier "medgen" ;
    rdfs:label "MedGen" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Mesh
    a owl:Class ;
    dcterms:identifier "mesh" ;
    rdfs:label "MeSH" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:MgiAllele
    a owl:Class ;
    dcterms:identifier "mgi_allele" ;
    rdfs:label "MGI allele" ;
    rdfs:subClassOf :Variant, :Dataset .

:MgiGene
    a owl:Class ;
    dcterms:identifier "mgi_gene" ;
    rdfs:label "MGI gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:MgiGenotype
    a owl:Class ;
    dcterms:identifier "mgi_genotype" ;
    rdfs:label "MGI genotype" ;
    rdfs:subClassOf :Genotype, :Dataset .

:Mirbase
    a owl:Class ;
    dcterms:identifier "mirbase" ;
    rdfs:label "miRBase" ;
    rdfs:subClassOf :Transcript, :Dataset .

:Mondo
    a owl:Class ;
    dcterms:identifier "mondo" ;
    rdfs:label "MONDO" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Mp
    a owl:Class ;
    dcterms:identifier "mp" ;
    rdfs:label "Mammalian Phenotype Ontology" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Nando
    a owl:Class ;
    dcterms:identifier "nando" ;
    rdfs:label "NANDO" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:NbdcHumanDb
    a owl:Class ;
    dcterms:identifier "nbdc_human_db" ;
    rdfs:label "NBDC Human Database" ;
    rdfs:subClassOf :Project, :Dataset .

:Ncbigene
    a owl:Class ;
    dcterms:identifier "ncbigene" ;
    rdfs:label "NCBI Gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:NcitDisease
    a owl:Class ;
    dcterms:identifier "ncit_disease" ;
    rdfs:label "NCIt disease" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:NcitTissue
    a owl:Class ;
    dcterms:identifier "ncit_tissue" ;
    rdfs:label "NCIt tissue" ;
    rdfs:subClassOf :Anatomy, :Dataset .

:OmaGroup
    a owl:Class ;
    dcterms:identifier "oma_group" ;
    rdfs:label "OMA group" ;
    rdfs:subClassOf :Ortholog, :Dataset .

:OmaProtein
    a owl:Class ;
    dcterms:identifier "oma_protein" ;
    rdfs:label "OMA protein" ;
    rdfs:subClassOf :Protein, :Dataset .

:OmimGene
    a owl:Class ;
    dcterms:identifier "omim_gene" ;
    rdfs:label "OMIM gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:OmimPhenotype
    a owl:Class ;
    dcterms:identifier "omim_phenotype" ;
    rdfs:label "OMIM phenotype" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:OrphanetGene
    a owl:Class ;
    dcterms:identifier "orphanet_gene" ;
    rdfs:label "Orphanet gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:OrphanetPhenotype
    a owl:Class ;
    dcterms:identifier "orphanet_phenotype" ;
    rdfs:label "Orphanet phenotype" ;
    rdfs:subClassOf :Phenotype, :Dataset .

:Pathbank
    a owl:Class ;
    dcterms:identifier "pathbank" ;
    rdfs:label "Pathbank" ;
    rdfs:subClassOf :Pathway, :Dataset .

:Pdb
    a owl:Class ;
    dcterms:identifier "pdb" ;
    rdfs:label "PDB" ;
    rdfs:subClassOf :Structure, :Dataset .

:PdbCcd
    a owl:Class ;
    dcterms:identifier "pdb_ccd" ;
    rdfs:label "PDB CCD" ;
    rdfs:subClassOf :Compound, :Dataset .

:Pfam
    a owl:Class ;
    dcterms:identifier "pfam" ;
    rdfs:label "Pfam" ;
    rdfs:subClassOf :Domain, :Dataset .

:Prosite
    a owl:Class ;
    dcterms:identifier "prosite" ;
    rdfs:label "PROSITE" ;
    rdfs:subClassOf :Domain, :Dataset .

:PrositeProrule
    a owl:Class ;
    dcterms:identifier "prosite_prorule" ;
    rdfs:label "PROSITE ProRule" ;
    rdfs:subClassOf :AnnotationRule, :Dataset .

:PubchemCompound
    a owl:Class ;
    dcterms:identifier "pubchem_compound" ;
    rdfs:label "PubChem compound" ;
    rdfs:subClassOf :Compound, :Dataset .

:PubchemPathway
    a owl:Class ;
    dcterms:identifier "pubchem_pathway" ;
    rdfs:label "PubChem pathway" ;
    rdfs:subClassOf :Pathway, :Dataset .

:PubchemSubstance
    a owl:Class ;
    dcterms:identifier "pubchem_substance" ;
    rdfs:label "PubChem substance" ;
    rdfs:subClassOf :Compound, :Dataset .

:Pubmed
    a owl:Class ;
    dcterms:identifier "pubmed" ;
    rdfs:label "PubMed" ;
    rdfs:subClassOf :Literature, :Dataset .

:ReactomePathway
    a owl:Class ;
    dcterms:identifier "reactome_pathway" ;
    rdfs:label "Reactome pathway" ;
    rdfs:subClassOf :Pathway, :Dataset .

:ReactomeReaction
    a owl:Class ;
    dcterms:identifier "reactome_reaction" ;
    rdfs:label "Reactome reaction" ;
    rdfs:subClassOf :Reaction, :Dataset .

:RefseqGenomic
    a owl:Class ;
    dcterms:identifier "refseq_genomic" ;
    rdfs:label "RefSeq genomic" ;
    rdfs:subClassOf :Gene, :Dataset .

:RefseqProtein
    a owl:Class ;
    dcterms:identifier "refseq_protein" ;
    rdfs:label "RefSeq protein" ;
    rdfs:subClassOf :Protein, :Dataset .

:RefseqRna
    a owl:Class ;
    dcterms:identifier "refseq_rna" ;
    rdfs:label "RefSeq RNA" ;
    rdfs:subClassOf :Transcript, :Dataset .

:Rgd
    a owl:Class ;
    dcterms:identifier "rgd" ;
    rdfs:label "RGD" ;
    rdfs:subClassOf :Gene, :Dataset .

:Rhea
    a owl:Class ;
    dcterms:identifier "rhea" ;
    rdfs:label "Rhea" ;
    rdfs:subClassOf :Reaction, :Dataset .

:Sgd
    a owl:Class ;
    dcterms:identifier "sgd" ;
    rdfs:label "SGD" ;
    rdfs:subClassOf :Gene, :Dataset .

:Smart
    a owl:Class ;
    dcterms:identifier "smart" ;
    rdfs:label "SMART" ;
    rdfs:subClassOf :Domain, :Dataset .

:SraAccession
    a owl:Class ;
    dcterms:identifier "sra_accession" ;
    rdfs:label "SRA accession" ;
    rdfs:subClassOf :Submission, :Dataset .

:SraAnalysis
    a owl:Class ;
    dcterms:identifier "sra_analysis" ;
    rdfs:label "SRA analysis" ;
    rdfs:subClassOf :Analysis, :Dataset .

:SraExperiment
    a owl:Class ;
    dcterms:identifier "sra_experiment" ;
    rdfs:label "SRA experiment" ;
    rdfs:subClassOf :Experiment, :Dataset .

:SraProject
    a owl:Class ;
    dcterms:identifier "sra_project" ;
    rdfs:label "SRA project" ;
    rdfs:subClassOf :Project, :Dataset .

:SraRun
    a owl:Class ;
    dcterms:identifier "sra_run" ;
    rdfs:label "SRA run" ;
    rdfs:subClassOf :SequenceRun, :Dataset .

:SraSample
    a owl:Class ;
    dcterms:identifier "sra_sample" ;
    rdfs:label "SRA sample" ;
    rdfs:subClassOf :Sample, :Dataset .

:Swisslipids
    a owl:Class ;
    dcterms:identifier "swisslipids" ;
    rdfs:label "SwissLipids" ;
    rdfs:subClassOf :Lipid, :Dataset .

:Tair
    a owl:Class ;
    dcterms:identifier "tair" ;
    rdfs:label "TAIR" ;
    rdfs:subClassOf :Gene, :Dataset .

:Taxonomy
    a owl:Class ;
    dcterms:identifier "taxonomy" ;
    rdfs:label "Taxonomy" ;
    rdfs:subClassOf :Organism, :Dataset .

:Togovar
    a owl:Class ;
    dcterms:identifier "togovar" ;
    rdfs:label "TogoVar variant" ;
    rdfs:subClassOf :Variant, :Dataset .

:Uberon
    a owl:Class ;
    dcterms:identifier "uberon" ;
    rdfs:label "UBERON" ;
    rdfs:subClassOf :Anatomy, :Dataset .

:Uniprot
    a owl:Class ;
    dcterms:identifier "uniprot" ;
    rdfs:label "UniProt" ;
    rdfs:subClassOf :Protein, :Dataset .

:UniprotMnemonic
    a owl:Class ;
    dcterms:identifier "uniprot_mnemonic" ;
    rdfs:label "UniProt mnemonic" ;
    rdfs:subClassOf :Protein , :Synonym, :Dataset .

:UniprotReferenceProteome
    a owl:Class ;
    dcterms:identifier "uniprot_reference_proteome" ;
    rdfs:label "UniProt reference proteome" ;
    rdfs:subClassOf :Proteome, :Dataset .

:Vgnc
    a owl:Class ;
    dcterms:identifier "vgnc" ;
    rdfs:label "VGNC" ;
    rdfs:subClassOf :Gene, :Dataset .

:Wikipathways
    a owl:Class ;
    dcterms:identifier "wikipathways" ;
    rdfs:label "WikiPathways" ;
    rdfs:subClassOf :Pathway, :Dataset .

:WormbaseGene
    a owl:Class ;
    dcterms:identifier "wormbase_gene" ;
    rdfs:label "WormBase gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:XenbaseGene
    a owl:Class ;
    dcterms:identifier "xenbase_gene" ;
    rdfs:label "Xenbase gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:ZfinGene
    a owl:Class ;
    dcterms:identifier "zfin_gene" ;
    rdfs:label "ZFIN gene" ;
    rdfs:subClassOf :Gene, :Dataset .

:TIO_000001
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Category ;
    rdfs:label "is nearly equivalent to" ;
    :display_label "is nearly equivalent to" .

:TIO_000002
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Category ;
    rdfs:label "is equivalent to" ;
    :display_label "is equivalent to" .

:TIO_000003
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Variant ;
    rdfs:range :Variant ;
    rdfs:label "variant has corresponding variant" ;
    :display_label "has corresponding variant" .

:TIO_000004
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Go ;
    owl:inverseOf :TIO_000007 ;
    rdfs:label "has GO annotation" ;
    :display_label "has GO annotation" .

:TIO_000005
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Go ;
    owl:inverseOf :TIO_000008 ;
    rdfs:label "has GO biological process" ;
    :display_label "has GO biological process" .

:TIO_000006
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Go ;
    owl:inverseOf :TIO_000009 ;
    rdfs:label "has GO cellular component" ;
    :display_label "has GO cellular component" .

:TIO_000007
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Go ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000004 ;
    rdfs:label "is GO annotation of" ;
    :display_label "is GO annotation of" .

:TIO_000008
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Go ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000005 ;
    rdfs:label "is GO biological process of" ;
    :display_label "is GO biological process of" .

:TIO_000009
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Go ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000006 ;
    rdfs:label "is GO cellular component of" ;
    :display_label "is GO cellular component of" .

:TIO_000010
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000011 ;
    rdfs:label "has related phenotype" ;
    :display_label "has related phenotype" .

:TIO_000011
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000010 ;
    rdfs:label "phenotype is related with" ;
    :display_label "is related with" .

:TIO_000012
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Molecule ;
    rdfs:range :Molecule ;
    rdfs:label "molecule interacts with molecule" ;
    :display_label "interacts with" .

:TIO_000013
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Domain ;
    owl:inverseOf :TIO_000015 ;
    rdfs:label "protein has protein domain" ;
    :display_label "has protein domain" .

:TIO_000014
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Structure ;
    rdfs:range :Domain ;
    owl:inverseOf :TIO_000016 ;
    rdfs:label "structure has protein domain" ;
    :display_label "has protein domain" .

:TIO_000015
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Domain ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000013 ;
    rdfs:label "protein domain is in protein" ;
    :display_label "is in protein" .

:TIO_000016
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Domain ;
    rdfs:range :Structure ;
    owl:inverseOf :TIO_000014 ;
    rdfs:label "protein domain is in structure" ;
    :display_label "is in structure" .

:TIO_000017
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Sequence ;
    rdfs:label "sequence is orthologous to sequence" ;
    :display_label "is orthologous to" .

:TIO_000018
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Ortholog ;
    rdfs:range :Sequence ;
    owl:inverseOf :TIO_000019 ;
    rdfs:label "orthologous group has member sequence" ;
    :display_label "has orthologous group member" .

:TIO_000019
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Ortholog ;
    owl:inverseOf :TIO_000018 ;
    rdfs:label "sequence is member of orthologous group" ;
    :display_label "is member of orthologous group" .

:TIO_000020
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Compound ;
    rdfs:range :InchiKey ;
    owl:inverseOf :TIO_000021 ;
    rdfs:label "compound is represented as InChIKey" ;
    :display_label "is represented as" .

:TIO_000021
    rdfs:subPropertyOf :relation ;
    rdfs:domain :InchiKey ;
    rdfs:range :Compound ;
    owl:inverseOf :TIO_000020 ;
    rdfs:label "InChIKey represents compound" ;
    :display_label "represents" .

:TIO_000022
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Synonym ;
    owl:inverseOf :TIO_000023 ;
    rdfs:label "has synonym" ;
    :display_label "has synonym" .

:TIO_000023
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Synonym ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000022 ;
    rdfs:label "is synonym of" ;
    :display_label "is synonym of" .

:TIO_000024
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000025 ;
    rdfs:label "protein is product of gene" ;
    :display_label "is product of gene" .

:TIO_000025
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000024 ;
    rdfs:label "gene has protein as product" ;
    :display_label "has gene product" .

:TIO_000026
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Structure ;
    owl:inverseOf :TIO_000027 ;
    rdfs:label "protein has 3D structure" ;
    :display_label "has 3D structure" .

:TIO_000027
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Structure ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000026 ;
    rdfs:label "is 3D structure of protein" ;
    :display_label "is 3D structure of" .

:TIO_000028
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Ec ;
    owl:inverseOf :TIO_000029 ;
    rdfs:label "sequence has EC number" ;
    :display_label "has EC number" .

:TIO_000029
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Ec ;
    rdfs:range :Sequence ;
    owl:inverseOf :TIO_000028 ;
    rdfs:label "is EC number of sequence" ;
    :display_label "is EC number of" .

:TIO_000030
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene, :Transcript ;
    rdfs:range :Probe ;
    owl:inverseOf :TIO_000031 ;
    rdfs:label "gene or transcript is target of probe" ;
    :display_label "is target of" .

:TIO_000031
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Probe ;
    rdfs:range :Gene, :Transcript ;
    owl:inverseOf :TIO_000030 ;
    rdfs:label "probe detects gene or transcript" ;
    :display_label "detects" .

:TIO_000032
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Literature ;
    owl:inverseOf :TIO_000033 ;
    rdfs:label "has reference" ;
    :display_label "has reference" .

:TIO_000033
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Literature ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000032 ;
    rdfs:label "is reference of" ;
    :display_label "is reference of" .

:TIO_000034
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Transcript ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000035 ;
    rdfs:label "transcript is transcribed from gene" ;
    :display_label "is transcribed from" .

:TIO_000035
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Transcript ;
    owl:inverseOf :TIO_000034 ;
    rdfs:label "gene is transcribed to transcript" ;
    :display_label "is transcribed to" .

:TIO_000036
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Transcript ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000037 ;
    rdfs:label "transcript is translated to protein" ;
    :display_label "is translated to" .

:TIO_000037
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Transcript ;
    owl:inverseOf :TIO_000036 ;
    rdfs:label "protein is translated from transcript" ;
    :display_label "is translated from" .

:TIO_000038
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Submission ;
    rdfs:range :Analysis ;
    owl:inverseOf :TIO_000048 ;
    rdfs:label "submission includes analysis" ;
    :display_label "includes analysis" .

:TIO_000039
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Submission ;
    rdfs:range :Experiment ;
    owl:inverseOf :TIO_000049 ;
    rdfs:label "submission includes experiment" ;
    :display_label "includes experiment" .

:TIO_000040
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :Experiment ;
    owl:inverseOf :TIO_000050 ;
    rdfs:label "project performs experiment" ;
    :display_label "performs experiment" .

:TIO_000041
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Submission ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000051 ;
    rdfs:label "submission includes project" ;
    :display_label "includes project" .

:TIO_000042
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Submission ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000052 ;
    rdfs:label "submission includes sample" ;
    :display_label "includes sample" .

:TIO_000043
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Experiment ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000053 ;
    rdfs:label "experiment captures sample" ;
    :display_label "captures sample" .

:TIO_000044
    rdfs:subPropertyOf :relation ;
    rdfs:domain :SequenceRun ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000054 ;
    rdfs:label "sequence run is produced from sample" ;
    :display_label "is produced from sample" .

:TIO_000045
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Submission ;
    rdfs:range :SequenceRun ;
    owl:inverseOf :TIO_000055 ;
    rdfs:label "submission includes sequence run" ;
    :display_label "includes sequence run" .

:TIO_000046
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :SequenceRun ;
    owl:inverseOf :TIO_000056 ;
    rdfs:label "project produces sequence run" ;
    :display_label "produces sequence run" .

:TIO_000047
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Experiment ;
    rdfs:range :SequenceRun ;
    owl:inverseOf :TIO_000057 ;
    rdfs:label "experiment produces sequence run" ;
    :display_label "produces sequence run" .

:TIO_000048
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Analysis ;
    rdfs:range :Submission ;
    owl:inverseOf :TIO_000038 ;
    rdfs:label "analysis belongs to submission" ;
    :display_label "belongs to submission" .

:TIO_000049
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Experiment ;
    rdfs:range :Submission ;
    owl:inverseOf :TIO_000039 ;
    rdfs:label "experiment belongs to submission" ;
    :display_label "belongs to submission" .

:TIO_000050
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Experiment ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000040 ;
    rdfs:label "experiment belongs to project" ;
    :display_label "belongs to project" .

:TIO_000051
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :Submission ;
    owl:inverseOf :TIO_000041 ;
    rdfs:label "project belongs to submission" ;
    :display_label "belongs to submission" .

:TIO_000052
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :Submission ;
    owl:inverseOf :TIO_000042 ;
    rdfs:label "sample belongs to submission" ;
    :display_label "belongs to submission" .

:TIO_000053
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :Experiment ;
    owl:inverseOf :TIO_000043 ;
    rdfs:label "sample is captured to perform experiment" ;
    :display_label "is captured to perform experiment" .

:TIO_000054
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :SequenceRun ;
    owl:inverseOf :TIO_000044 ;
    rdfs:label "sample is captured to produce sequence run" ;
    :display_label "is captured to produce sequence run" .

:TIO_000055
    rdfs:subPropertyOf :relation ;
    rdfs:domain :SequenceRun ;
    rdfs:range :Submission ;
    owl:inverseOf :TIO_000045 ;
    rdfs:label "sequence run belongs to submission" ;
    :display_label "belongs to submission" .

:TIO_000056
    rdfs:subPropertyOf :relation ;
    rdfs:domain :SequenceRun ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000046 ;
    rdfs:label "sequence run belongs to project" ;
    :display_label "belongs to project" .

:TIO_000057
    rdfs:subPropertyOf :relation ;
    rdfs:domain :SequenceRun ;
    rdfs:range :Experiment ;
    owl:inverseOf :TIO_000047 ;
    rdfs:label "sequence run is produced in experiment" ;
    :display_label "is produced in experiment" .

:TIO_000058
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Variant ;
    rdfs:range :Variant ;
    owl:inverseOf :TIO_000059 ;
    rdfs:label "variant has clinical significance" ;
    :display_label "has clinical significance" .

:TIO_000059
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Variant ;
    rdfs:range :Variant ;
    owl:inverseOf :TIO_000058 ;
    rdfs:label "is clinical significance of" ;
    :display_label "is clinical significance of" .

:TIO_000060
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Glycan ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000061 ;
    rdfs:label "glycan is attached to protein" ;
    :display_label "is attached to" .

:TIO_000061
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Glycan ;
    owl:inverseOf :TIO_000060 ;
    rdfs:label "protein is modified with glycan" ;
    :display_label "is modified with" .

:TIO_000062
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Variant ;
    rdfs:range :Sequence ;
    owl:inverseOf :TIO_000063 ;
    rdfs:label "variant is located in sequence" ;
    :display_label "is located in" .

:TIO_000063
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Variant ;
    owl:inverseOf :TIO_000062 ;
    rdfs:label "sequence has variant" ;
    :display_label "has variant" .

:TIO_000064
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Organism ;
    owl:inverseOf :TIO_000065 ;
    rdfs:label "is gene of organism" ;
    :display_label "is gene of organism" .

:TIO_000065
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Organism ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000064 ;
    rdfs:label "organism has gene" ;
    :display_label "has gene" .

:TIO_000066
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Compound ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000067 ;
    rdfs:label "compound targets to phenotype" ;
    :display_label "targets to" .

:TIO_000067
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Compound ;
    owl:inverseOf :TIO_000066 ;
    rdfs:label "phenotype is targeted by compound" ;
    :display_label "is targeted by" .

:TIO_000068
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Pathway ;
    rdfs:range :Molecule ;
    owl:inverseOf :TIO_000069 ;
    rdfs:label "pathway has participant molecule" ;
    :display_label "has participant molecule" .

:TIO_000069
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Molecule ;
    rdfs:range :Pathway ;
    owl:inverseOf :TIO_000068 ;
    rdfs:label "molecule participates in pathway" ;
    :display_label "participates in pathway" .

:TIO_000070
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Pathway ;
    rdfs:range :Reaction ;
    owl:inverseOf :TIO_000071 ;
    rdfs:label "pathway has participant reaction" ;
    :display_label "has participant reaction" .

:TIO_000071
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Reaction ;
    rdfs:range :Pathway ;
    owl:inverseOf :TIO_000070 ;
    rdfs:label "reaction participates in pathway" ;
    :display_label "participates in pathway" .

:TIO_000072
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Pathway ;
    rdfs:range :Domain ;
    owl:inverseOf :TIO_000073 ;
    rdfs:label "pathway has protein domain" ;
    :display_label "has protein domain in pathway" .

:TIO_000073
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Domain ;
    rdfs:range :Pathway ;
    owl:inverseOf :TIO_000072 ;
    rdfs:label "protein domain participates in pathway" ;
    :display_label "participates in pathway" .

:TIO_000074
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Molecule ;
    rdfs:range :Interaction ;
    owl:inverseOf :TIO_000075 ;
    rdfs:label "molecule participates in interaction" ;
    :display_label "participates in interaction" .

:TIO_000075
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Interaction ;
    rdfs:range :Molecule ;
    owl:inverseOf :TIO_000074 ;
    rdfs:label "interaction has participant molecule" ;
    :display_label "has participant molecule" .

:TIO_000076
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Molecule ;
    rdfs:range :Reaction ;
    owl:inverseOf :TIO_000077 ;
    rdfs:label "molecule participates in reaction" ;
    :display_label "participates in reaction" .

:TIO_000077
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Reaction ;
    rdfs:range :Molecule ;
    owl:inverseOf :TIO_000076 ;
    rdfs:label "reaction has participant molecule" ;
    :display_label "has participant molecule" .

:TIO_000078
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Genome ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000079 ;
    rdfs:label "genome is sequenced in project" ;
    :display_label "is sequenced in" .

:TIO_000079
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :Genome ;
    owl:inverseOf :TIO_000078 ;
    rdfs:label "project determines genome sequence" ;
    :display_label "determines genome sequence" .

:TIO_000080
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Genome ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000081 ;
    rdfs:label "genome is sequenced with sample" ;
    :display_label "is sequenced with" .

:TIO_000081
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :Genome ;
    owl:inverseOf :TIO_000080 ;
    rdfs:label "sample is used to sequence genome" ;
    :display_label "is used to sequence" .

:TIO_000082
    rdfs:subPropertyOf :relation ;
    rdfs:domain :CellLine ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000083 ;
    rdfs:label "cell line is model of phenotype" ;
    :display_label "is model of" .

:TIO_000083
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :CellLine ;
    owl:inverseOf :TIO_000082 ;
    rdfs:label "phenotype is observed in cell line" ;
    :display_label "is observed in" .

:TIO_000084
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Pathway ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000085 ;
    rdfs:label "pathway has participant gene" ;
    :display_label "has participant gene" .

:TIO_000085
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Pathway ;
    owl:inverseOf :TIO_000084 ;
    rdfs:label "gene participates in pathway" ;
    :display_label "participates in pathway" .

:TIO_000086
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Anatomy ;
    owl:inverseOf :TIO_000087 ;
    rdfs:label "phenotype is observed in anatomic site" ;
    :display_label "is observed in anatomic site" .

:TIO_000087
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Anatomy ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000086 ;
    rdfs:label "anatomic site can have phenotype" ;
    :display_label "can have phenotype" .

:TIO_000088
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Drug ;
    rdfs:range :Compound ;
    owl:inverseOf :TIO_000089 ;
    rdfs:label "drug classification classifies compound" ;
    :display_label "classifies" .

:TIO_000089
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Compound ;
    rdfs:range :Drug ;
    owl:inverseOf :TIO_000088 ;
    rdfs:label "compound is classified as drug classification" ;
    :display_label "is classified as" .

:TIO_000090
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Proteome ;
    rdfs:range :Organism ;
    owl:inverseOf :TIO_000091 ;
    rdfs:label "is proteome of organism" ;
    :display_label "is proteome of" .

:TIO_000091
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Organism ;
    rdfs:range :Proteome ;
    owl:inverseOf :TIO_000090 ;
    rdfs:label "organism has proteome" ;
    :display_label "has proteome" .

:TIO_000092
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Proteome ;
    rdfs:range :Genome ;
    owl:inverseOf :TIO_000093 ;
    rdfs:label "proteome is coded by genome" ;
    :display_label "is coded by" .

:TIO_000093
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Genome ;
    rdfs:range :Proteome ;
    owl:inverseOf :TIO_000092 ;
    rdfs:label "genome codes proteome" ;
    :display_label "codes" .

:TIO_000094
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000095 ;
    rdfs:label "project analyzes sample" ;
    :display_label "analyzes" .

:TIO_000095
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000094 ;
    rdfs:label "sample is analyzed in project" ;
    :display_label "is analyzed in" .

:TIO_000096
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Structure ;
    rdfs:range :Compound ;
    owl:inverseOf :TIO_000097 ;
    rdfs:label "3D structure has compound" ;
    :display_label "has compound" .

:TIO_000097
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Compound ;
    rdfs:range :Structure ;
    owl:inverseOf :TIO_000096 ;
    rdfs:label "compound is component of 3D structure" ;
    :display_label "is component of" .

:TIO_000098
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Glycan ;
    owl:inverseOf :TIO_000099 ;
    rdfs:label "gene codes protein to process glycan" ;
    :display_label "codes protein to process glycan" .

:TIO_000099
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Glycan ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000098 ;
    rdfs:label "glycan is processed by protein coded by gene" ;
    :display_label "is processed by protein coded by gene" .

:TIO_000100
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000101 ;
    rdfs:label "disease has symptom" ;
    :display_label "has symptom" .

:TIO_000101
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000100 ;
    rdfs:label "symptom is observed in disease" ;
    :display_label "is observed in disease" .

:TIO_000102
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Symptom ;
    owl:inverseOf :TIO_000103 ;
    rdfs:label "gene is related with symptom" ;
    :display_label "is related with symptom" .

:TIO_000103
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Symptom ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000102 ;
    rdfs:label "symptom is related with gene" ;
    :display_label "is related with gene" .

:TIO_000104
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Phenotype ;
    rdfs:range :Classification ;
    owl:inverseOf :TIO_000105 ;
    rdfs:label "phenotype has mode of inheritance" ;
    :display_label "has mode of inferitance" .

:TIO_000105
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Classification ;
    rdfs:range :Phenotype ;
    owl:inverseOf :TIO_000104 ;
    rdfs:label "is mode of inheritance of phenotype" ;
    :display_label "is mode of inheritance of" .

:TIO_000106
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000107 ;
    rdfs:label "has member" ;
    :display_label "has member" .

:TIO_000107
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Category ;
    rdfs:range :Category ;
    owl:inverseOf :TIO_000106 ;
    rdfs:label "is member of" ;
    :display_label "is member of" .

:TIO_000108
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Project ;
    owl:inverseOf :TIO_000109 ;
    rdfs:label "sequence is determined in project" ;
    :display_label "is determined in project" .

:TIO_000109
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Project ;
    rdfs:range :Sequence ;
    owl:inverseOf :TIO_000108 ;
    rdfs:label "project determines sequence" ;
    :display_label "determines sequence" .

:TIO_000110
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sequence ;
    rdfs:range :Sample ;
    owl:inverseOf :TIO_000111 ;
    rdfs:label "sequence is determined using sample" ;
    :display_label "is determined using sample" .

:TIO_000111
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Sample ;
    rdfs:range :Sequence ;
    owl:inverseOf :TIO_000110 ;
    rdfs:label "sample is used to determine sequence" ;
    :display_label "is used to determine sequence" .

:TIO_000112
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Gene ;
    rdfs:range :Pathway ;
    owl:inverseOf :TIO_000113 ;
    rdfs:label "gene codes protein participating in pathway" ;
    :display_label "codes protein participating in" .

:TIO_000113
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Pathway ;
    rdfs:range :Gene ;
    owl:inverseOf :TIO_000112 ;
    rdfs:label "pathway has component coded by gene" ;
    :display_label "has component coded by" .

:TIO_000114
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Domain ;
    rdfs:range :Function ;
    owl:inverseOf :TIO_000115 ;
    rdfs:label "domain has annotation rule" ;
    :display_label "has annotation rule" .

:TIO_000115
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Function ;
    rdfs:range :Domain ;
    owl:inverseOf :TIO_000114 ;
    rdfs:label "is annotation rule of" ;
    :display_label "is annotation rule of" .

:TIO_000116
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Genotype ;
    rdfs:range :Variant ;
    owl:inverseOf :TIO_000117 ;
    rdfs:label "genotype is composed of allele" ;
    :display_label "is composed of allele" .

:TIO_000117
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Variant ;
    rdfs:range :Genotype ;
    owl:inverseOf :TIO_000116 ;
    rdfs:label "allele composes genotype" ;
    :display_label "composes genotype" .

:TIO_000118
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Protein ;
    rdfs:range :Glycan ;
    owl:inverseOf :TIO_000119 ;
    rdfs:label "protein processes glycan" ;
    :display_label "processes" .

:TIO_000119
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Glycan ;
    rdfs:range :Protein ;
    owl:inverseOf :TIO_000118 ;
    rdfs:label "glycan is processed by protein" ;
    :display_label "is processed by" .

:TIO_000120
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Compound ;
    rdfs:range :Molecule ;
    owl:inverseOf :TIO_000121 ;
    rdfs:label "compound targets to molecule" ;
    :display_label "targets to" .

:TIO_000121
    rdfs:subPropertyOf :relation ;
    rdfs:domain :Molecule ;
    rdfs:range :Compound ;
    owl:inverseOf :TIO_000120 ;
    rdfs:label "molecule is targeted by compound" ;
    :display_label "is targeted by" .